The Ancient Life History of the Earth: A Comprehensive Outline of the Principles and Leading Facts of Palæontological ScienceNicholson, Henry Alleyne
Science
The Ancient Life History of the Earth: A Comprehensive Outline of the Principles and Leading Facts of Palæontological Science
Nicholson, Henry Alleyne
Paleontology
Finally, we meet in the Permian deposits with the first undoubted
remains of true _Reptiles_. These are distinguished, as a class,
from the _Amphibians_, by the fact that they are air-breathers
throughout the whole of their life, and therefore are at no time
provided with gills; whilst they are exempt from that metamorphosis
which all the _Amphibia_ undergo in early life, consequent upon
their transition from an aquatic to a more or less purely aerial
mode of respiration. Their skeleton is well ossified; they usually
have horny or bony plates, singly or in combination, developed
in the skin; and their limbs (when present) are never either
in the form of _fins_ or _wings_, though sometimes capable of
acting in either of these capacities, and liable to great
modifications of form and structure. Though there can be no doubt
whatever as to the occurrence of genuine Reptiles in deposits of
unquestionable Permian age, there is still uncertainty as to the
precise number of types which may have existed at this period.
This uncertainty arises partly from the difficulty of deciding
in all cases, whether a given bone be truely Labyrinthodont or
Reptilian, but more especially from the confusion which exists at
present between the Permian and the overlying Triassic deposits.
Thus there are various deposits in different regions which have
yielded the remains of Reptiles, and which cannot in the meanwhile
be definitely referred either to the Permian series or to the
Trias by clear stratigraphical or palæontological evidence. All
that can be done in such cases is to be guided by the characters
of the Reptiles themselves, and to judge by their affinities to
remains from known Triassic or Permian rocks to which of these
formations the beds containing them should be referred; but it
is obvious that this method of procedure is seriously liable
to lead to error. In accordance, however, with this, the only
available mode of determination in some cases, the remains of
_Thecodontosaurus_ and _Palæosaurus_ discovered in the dolomitic
conglomerates near Bristol will be considered as Triassic, thus
leaving _Protorosaurus_[20] as the principal and most important
representative of the Permian Reptiles.[21] The type-species of
the genus _Protorusaurus_ is the _P. Speneri_(fig. 138) of the
"Kupfer-schiefer" of Thuringia, but other allied species have
been detected in the Middle Permian of Germany and the north
of England. This Reptile attained a length of from three to four
feet; and it has been generally referred to the group of the
Lizards (_Lacertilia_), to which it is most nearly allied in
its general structure, at the same time that it differs from
all existing members of this group in the fact that its numerous
conical and pointed teeth were implanted in distinct sockets in
the jaws--this being a Crocodilian character. In other respects,
however, _Protorosaurus_ approximates closely to the living Monitors
(_Varanidoe_); and the fact that the bodies of the vertebræ are
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