The Ancient Life History of the Earth: A Comprehensive Outline of the Principles and Leading Facts of Palæontological ScienceNicholson, Henry Alleyne
Science
The Ancient Life History of the Earth: A Comprehensive Outline of the Principles and Leading Facts of Palæontological Science
Nicholson, Henry Alleyne
Paleontology
Amongst the _Vertebrate Animals_ of the Trias, the _Fishes_ are
represented by numerous forms belonging to the _Ganoids_ and the
_Placoids_. The Ganoids of the period are still all provided
with unsymmetrical ("heterocercal") tails, and belong principally
to such genera as _Paloeoniscus_ and _Catopterus_. The remains of
Placoids are in the form of teeth and spines, the two principal
genera being the two important Secondary groups _Acrodus_ and
_Hybodus_. Very nearly at the summit of the Trias in England, in
the Rhætic series, is a singular stratum, which is well known as the
"bone-bed," from the number of fish-remains which it contains. More
interesting, however, than the above, are the curious palate-teeth
of the Trias, upon which Agassiz founded the genus _Ceratodus_.
The teeth of Ceratodus (fig. 146) are singular flattened plates,
composed of spongy bone beneath, covered superficially with a
layer of enamel. Each plate is approximately triangular, one
margin (which we now know to be the outer one) being prolonged
into prongs or conical prominences, whilst the surface is more or
less regularly undulated. Until recently, though the master-mind
of Agassiz recognised that these singular bodies were undoubtedly
the teeth of fishes, we were entirely ignorant as to their precise
relation to the animal, or as to the exact affinities of the fish
thus armed. Lately, however, there has been discovered in the
rivers of Queensland (Australia) a living species of _Ceratodus_
(_C. Fosteri_, fig. 147), with teeth precisely similar to those
of its Triassic predecessor; and we thus have become acquainted
with the use of these structures and the manner in which they
were implanted in the mouth. The palate carries two of these
plates, with their longer straight sides turned towards each
other, their sharply-sinuated sides turned outwards, and their
short straight sides or bases directed backwards. Two similar
plates in the lower jaw correspond to the upper, their undulated
surfaces fitting exactly to those of the opposite teeth. There
are also two sharp-edged front teeth, which are placed in the
front of the mouth in the upper jaw; but these have not been
recognised in the fossil specimens. The living _Ceratodus_ feeds
on vegetable matters, which are taken up or tom off from plants
by the sharp front teeth, and then partially crushed between
the undulated surfaces of the back teeth (Günther); and there
need be little doubt but that the Triassic _Ceratodi_ followed
a similar mode of existence. From the study of the living
_Ceratodus_, it is certain that the genus belongs to the same
group as the existing Mud-fishes (_Dipnoi_); and we therefore
learn that this, the highest, group of the entire class of Fishes
existed in Triassic times under forms little or not at all different
from species now alive; whilst it has become probable that the
order can be traced back into the Devonian period.
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