The Ancient Life History of the Earth: A Comprehensive Outline of the Principles and Leading Facts of Palæontological ScienceNicholson, Henry Alleyne
Science
The Ancient Life History of the Earth: A Comprehensive Outline of the Principles and Leading Facts of Palæontological Science
Nicholson, Henry Alleyne
Paleontology
The great class of the Birds, as we have seen, is represented
in rocks earlier than the Oolites simply by the not absolutely
certain evidence of the three-toed footprints of the Connecticut
Trias. In the Lithographic Slate of Solenhofen (Middle Oolite),
there has been discovered, however, the at present unique skeleton
of a Bird well known under the name of the _Archoeopteryx macrura_
(figs. 181, 182). The only known specimen--now in the British
Museum--unfortunately does not exhibit the skull; but the
fine-grained matrix has preserved a number of the other bones
of the skeleton, along with the impressions of the tail and wing
feathers. From these remains we know that _Archoeopteryx_ differed
in some remarkable peculiarities of its structure from all existing
members of the class of Birds. This extraordinary Bird (fig.
182) appears to have been about as big as a Rook--the tail being
long and extremely slender, and composed of separate vertebræ,
each of which supports a single pair of quill-feathers. In the
flying Birds of the present day, as before mentioned, the terminal
vertebræ of the tail are amalgamated to form a single bone
("ploughshare-bone"), which supports a cluster of tail-feathers;
and the tail itself is short. In the embryos of existing Birds
the tail is long, and is made up of separate vertebræ, and the
same character is observed in many existing Reptiles. The tail
of _Archoeopteryx_, therefore, is to be regarded as the permanent
retention of an embryonic type of structure, or as an approximation
to the characters of the Reptiles. Another remarkable point in
connection with _Archoeopteryx_, in which it differs from all
known Birds, is, that the wing was furnished with two free claws.
From the presence of feathers, _Archoeopteryx_ may be inferred to
have been hot-blooded; and this character, taken along with the
structure of the skeleton of the wing, may be held as sufficient
to justify its being considered as belonging to the class of
Birds. In the structure of the tail, however, it is singularly
Reptilian; and there is reason to believe that its jaws were
furnished with teeth sunk in distinct sockets, as is the case
in no existing Bird. This conclusion, at any rate, is rendered
highly probable by the recent discovery of "Toothed Birds"
(_Odonturnithes_) in the Cretaceous rocks of North America.
[Illustration: Fig. 183.--Lower jaw of _Amphitherium_
(_Thylacotherium_) _Prevostii_. Stonesfield Slate (Great Oolite.)]
[Illustration: Fig. 184. Oolitic Mammals.--1, Lower jaw and teeth
of _Phascolotherium_, Stonesfield Slate; 2, Lower jaw and teeth
of _Amphitherium_, Stonesfield Slate; 3, Lower jaw and teeth of
_Triconodon_, Purbeck beds; 4, Lower jaw and teeth of _Plagiaulax_,
Purbeck beds. All the figures are of the natural size.]
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