The Animal Parasites of ManFantham, Harold Benjamin
Science
The Animal Parasites of Man
Fantham, Harold Benjamin
Medical parasitology
The peripheral nerves, which spring from the posterior strands as well
as from the commissures, either pass directly to the muscular fibres or
to the sensory cells that are situated at the level of the subcuticular
cells, or they reach these after the formation of a plexus situated
immediately beneath the dermo-muscular layer; the processes directed
outwards terminate in small vesicles in the cuticle.
As to other ORGANS OF SENSE, simple eyes, two or four in number, are
known in several ectoparasitic species as well as in a few free-living
larval stages (Cercariæ) of endoparasitic forms. In the adult stage,
however, they usually undergo complete atrophy.
The ALIMENTARY CANAL commences with an oral aperture, generally
terminal or sub-terminal (ventral) at the anterior extremity, which
leads into an oral cavity usually surrounded by a sucker; the
œsophagus, of various lengths, is directed backwards and is generally
surrounded by a muscular pharynx (fig. 122). In some cases there exists
between the sucker and pharynx, pharyngeal pouches (præpharynx). Sooner
or later the intestine divides into two lateral branches directed
backwards, both of which end blindly (cæca) at the same level.[257]
In many ectoparasites (_Monogenea_ [p. 222]) a connection exists
between the genital glands and one of the intestinal branches (ductus
vitello-intestinalis [fig. 123]).
[257] The following conditions represent deviations from this type: (1)
In _Gasterostomum_ the oral aperture is situated in the middle of the
ventral surface, and occasionally is even nearer to the posterior than
to the anterior end. There is no proper oral sucker, but the pharynx is
thus termed. (2) A few genera, such as _Gasterostomum_, _Aspidogaster_,
_Diplozoon_, etc., have only _one_ intestinal diverticulum, which is
undoubtedly to be taken as representing the primitive condition, as
it is also often found in the young stages of the _Trematoda_. (3)
The branches of the intestines are curved and united behind (several
_Tristomidæ_ and _Monostomidæ_), while in _Polystomum integerrimum_
(in the bladder of frogs) there are several commissures between the
intestinal branches, and in the _Schistosomidæ_ the united intestinal
branches proceed as one channel towards the posterior end. (4) The
termination of the two intestinal branches is not always on a level;
they are therefore of different lengths. (5) When the œsophagus is
very long the intestinal branches extend both forward and backward, so
that the gut exhibits the form of an *H*. (6) In the broad and flat
species the gut-forks form diverticula mostly externally but also
internally; these again may branch (fig. 139). (7) In a few cases
(_Nematobothrium_, _Didymozoon_) the intestine completely disappears up
to the pharynx.
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