The Animal Parasites of ManFantham, Harold Benjamin
Science
The Animal Parasites of Man
Fantham, Harold Benjamin
Medical parasitology
The EXCRETORY SYSTEM[258] consists of several parts: (1) of the more or
less numerous terminal “flame” cells or funnel cells (figs. 124, 125);
(2) of the capillaries ending in them; (3) of larger vessels receiving
the capillaries; and (4) of the excretory bladder. Terminal cells and
capillaries may be compared to unicellular glands with long excretory
ducts; the cellular body (fig. 125) is comparatively large, stretched
longitudinally, more rarely transversely, and provided with numerous
processes, that are lost in the parenchyma; within is a conical cavity
(analogous to the secretory cavity of unicellular glands) which is
continued directly into the structureless capillary; at its blind end
is a bunch of cilia projecting into the cavity, and which, during life,
shows a flickering motion (ciliary flame). The nucleus is situated in
the protoplasm of the terminal cell at its blind end.
[258] The following description relates in the main to the _Distomata_.
The entire apparatus thus begins blindly--_i.e._, within the terminal
cells, to which must be ascribed the capacity of taking up from the
fluid that permeates the parenchyma the products which are first
collected into their own cavities and thence excreted by means of the
capillaries and vessels.
[Illustration: FIG. 125.--Terminal flame cell of the excretory system.
_n._, nucleus of cell; _c._, bundle of cilia forming the “flame”; _p._,
processes of cell extending into parenchyma; _d._, excretory capillary.
(Stephens.)]
The vessels possess definite walls, consisting of a membrane and a
nucleated protoplasmic layer. They unite at many points on either side,
and again pass into other canals (COLLECTING TUBES), which finally,
travelling towards the posterior end, discharge into the excretory
bladder (fig. 124).
The form and size of the bladder vary much according to the different
species, but it always possesses its own flattened epithelium,
surrounded by circular and longitudinal muscles, the circular muscles
forming a sphincter around the opening. Frequently also the structure
of the bladder extends to the tubules discharging into it, which
therefore are not to be regarded as separate “vessels,” but rather
as tubular diverticula of the bladder, directed anteriorly. In some
few species the diverticula also branch and the branches anastomose,
so that a network of tubules ensues which receives the vessels or
capillaries. In such cases there are also ciliary tracts in the tubules.
The contents of the entire apparatus usually consist of a clear or
sometimes reddish fluid; in some species there are larger or smaller
granules, and occasionally also concretions occur.
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