The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
Burmeister (1843; 1846, p. 19) considered the two kinds of eyes as
essentially the same, and accounted for the conspicuous lenses of
Phacops on the supposition that the cornea was thinner in that genus
than in the trilobites with smooth eyes.
Barrande (1852, p. 135) recognized three types of eyes in trilobites,
adding to Quenstedt's smooth and facetted compound eyes the groups of
simple eyes found in Harpes. In his sections of 1852, pl. 3, figs.
15-25, which are evidently diagrammatic, he shows separated biconvex
lenses in both types of compound eyes, _Phacops_ and _Dalmanites_ on
one hand, and _Asaphus_, _Goldius_, _Acidaspis_, and _Cyclopyge_
on the other. Clarke ( 1888), Exner ( 1891 ) and especially
Lindstroem (1901) have since published much more accurate figures and
descriptions. The first person to study the eye in thin section seems
to have been Packard (1880), who published some very sketchy figures
of specimens loaned him by Walcott. He studied the eyes of _Isotelus
gigas_, _Bathyurus longispinus_, _Calymene_, and _Phacops_, and
decided that the two types of eyes were fundamentally the same.
He also compared them with the eyes of _Limulus_.
Clarke (1888), in a careful study of the eye of _Phacops rana_, found
that the lenses were unequally biconvex, the curvature greater on the
inner surface. The lens had a circular opening on the inner side,
leading into a small pear-shaped cavity. The individual lenses were
quite distinct from one another, and separated by a continuation of
the test of the cheek.
Exner (1891, p. 34), in a comparison of the eyes of Phacops and
_Limulus_, came to the opinion that they were very unlike, and that
the former were really aggregates of simple eyes.
Lindstroem (1901, pp. 27-31) came to the conclusion that besides the
blind trilobites there were trilobites with two kinds of compound
eyes, trilobites with aggregate eyes, and trilobites with stemmata and
ocelli. His views may be briefly summarized.
I. Compound eyes.
1. Eyes with prismatic, plano-convex lenses.
"A pellucid, smooth and glossy integument, a direct continuation of
the common test of the body, covers the corneal lenses, quite as is
the case in so many of the recent Crustacea. The lenses are closely
packed, minute, usually hexagonal in outline, flat on the outer and
convex on the inner surface. Such eyes are best developed in
_Asaphus_, _Illænus_, _Nileus_, _Bumastus_, _Proëtus_, etc."
2. Eyes with biconvex lenses.
The surface of the eye is a mass of contiguous lenses, covered by a
thin membrane which is frequently absent from the specimens, due to
poor preservation. The lenses are biconvex, and being in contact
with one another, are usually hexagonal, although in some cases
they nearly retain their globular shape. Such eyes are found in
Bury care, _Peltura_, _Sphæropthalmus_, _Ctenopyge_, _Goldius_,
_Cheirurus_, and probably others.
II. Aggregate eyes.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account