The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
The oldest known isopod is _Oxyuropoda ligioides_ Carpenter and
Swain (Proc. Royal Irish Acad., vol. 27, sect. B, 1908, p. 63,
fig. 1), found in the Upper Devonian of County Kilkenny, Ireland. The
appendages are not known, but the test is in some ways like that of a
trilobite. The thorax, abdomen, and pygidium are especially like those
of certain trilobites, and there is no greater differentiation between
thorax and abdomen than there is between the regions before and behind
the fifteenth segment of a _Pædeumias_ or _Mesonacis_. The anal
segment is directly comparable to the pygidium of a _Ceraurus_, the
stiff unsegmented uropods being like the great lateral spines of that
genus.
The interpretation of the head offered by Carpenter and Swain is very
difficult to understand, as their description and figure do not seem
to agree. What they consider the first thoracic segment (fused with
the head) seems to me to be the posterior part of the cephalon and it
shows at the back a narrow transverse area which is at least analogous
to the nuchal segment of the trilobite. If this interpretation can be
sustained, _Oxyuropoda_ would be a very primitive isopod in which the
first thoracic segment (second of Carpenter and Swain) is still free.
According to the interpretation of the original authors, the species
is more specialized than recent Isopoda, as they claim that two
thoracic segments are fused in the head. The second interpretation was
perhaps made on the basis of the number of segments (nineteen) in a
recent isopod.
=Marrella splendens= WALCOTT.
Illustrated: Walcott, Smithson. Misc. Coll., vol. 57, 1912, p. 192,
pls. 25, 26.
Among the most wonderful of the specimens described by Doctor
Walcott is the "lace crab." While the systematic position was not
satisfactorily determined by the describer, it has been aptly compared
to a trilobite. The great nuchal and genal spines and the large
marginal sessile eyes, coupled with the almost total lack of thoracic
and abdominal test, give it a bizarre appearance which may obscure its
real relationships.
The cephalon appears to bear five pairs of appendages, antennules, and
antennæ, both tactile organs with numerous short segments, mandibles,
and first and second maxillæ. The last three pairs are elongate, very
spinose limbs, of peculiar appearance. They seem to have seven
segments, but are not well preserved. These organs are attached near
the posterior end of the labrum.
There are twenty-four pairs of biramous thoracic appendages, which
lack endobases. The endopodites are long and slender, with numerous
spines; the exopodites have narrow, thin shafts, with long, forward
pointed setæ. The anal segment consists of a single plate.
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