The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
The specialization of _Sidneyia_ consists in the remarkable
development of a highly complex claw on each of the third cephalic
appendages, and in the compound tail-fin, built up of the last segment
and one or more pairs of swimmerets. These two characteristics seem to
preclude the possibility of deriving the eurypterids from _Sidneyia_
itself, but it seems entirely within reason that they may have been
derived from another slightly less specialized member of the same
order.
That _Sidneyia_ is descended from any known trilobite seems highly
improbable, but that it was descended from the same ancestral stock as
the trilobites is, I believe, indicated by the presence of five pairs
of appendages on the cephalon and trilobitic legs on the abdomen.
=Molaria= and =Habelia.=
Other so-called Merostomata found by Walcott in the Middle Cambrian
are the genera _Molaria_ and _Habelia_, both referred to the Cambrian
family Aglaspidæ. These genera seem to conform with _Aglaspis_ of the
Upper Cambrian in having a trilobite-like cephalon without facial
sutures, a trilobite-like thorax of a small but variable (7-12) number
of segments, and a _Limulus_-like telson. Neither of them has yet been
fully described or figured, but (Walcott 1912 A, p. 202) _Habelia_
appears to have five pairs of cephalic appendages, the first two pairs
of which are multisegmented antennæ. The thoracic appendages are
likewise none too well known, but they appear to have been biramous.
The endopodites are better preserved than the exopodites, but in at
least one specimen of _Molaria_ the exopodites are conspicuous.
If these genera are properly described and figured, their appendages
are typically crustacean, and fundamentally in agreement with those of
_Marrella_. The relation to the Trilobita is evidently close, the
principal differences being the absence of facial sutures and the
presence of true antennæ. I am therefore transferring the Aglaspidæ
from the Merostomata to a new subclass under the Crustacea.
ARANEÆ.
The spiders have the head and thorax fused, the abdomen unsegmented
except in the most primitive suborder, and so appear even less
trilobite-like than the insects. The appendages likewise are highly
specialized. The cephalothorax bears six pairs of appendages, the
first of which are the pre-oral cheliceræ, while behind the mouth are
the pedipalpi and four pairs of ambulatory legs. The posterior pairs
of walking legs belong to the thorax, but the anterior ones are to be
homologized with the maxillæ of Crustacea, so that the spiders are
like the trilobites in having functional walking legs on the head.
Public-domain text, read in full here on John Shaqi.
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