The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
From the phylogeny of certain groups, such as the Asaphidæ, it is
learned that the geologically older members of the family have more
strongly segmented anterior and posterior shields than the later ones.
That there has been a "smoothing out" is demonstrated by a study of
the ontogeny of the later forms. From such examples it has come to
be thought that all smooth trilobites are specialized and occupy a
terminal position in their genealogical line. This has caused some
wonder that smooth agnostids like _Phalacroma bibullatum_ and _P.
nudum_ should be found in strata so old as the Middle Cambrian, and
was a source of great perplexity to me in the case of _Weymouthia_
(Ottawa Nat., vol. 27, 1913) (fig. 35). This is a smooth member of the
Eodiscidæ, and, in fact, one of the simplest trilobites known, for
while it has three thoracic segments, it shows almost no trace of
dorsal furrows or segmentation on cephalon or pygidium, and, of
course, no eyes. Following the general rule, I took this to be a
smooth-out eodiscid, and was surprised that it should come from the
Lower Cambrian, where it is associated with _Elliptocephala_ at Troy,
New York, and with _Callavia_ at North Weymouth, Massachusetts, and
where it has lately been found by Kiær associated with _Holmia_ and
_Kjerulfia_ at Tømten, Norway. It now appears it is really in its
proper zone, and instead of being the most specialized, is the
simplest of the Eodiscidæ.
What appears to be a still simpler trilobite is the form described by
Walcott as Naraoia.
=Naraoia compacta= Walcott.
(Text fig. 36.)
Illustrated: Walcott, Smithson. Misc. Coll., vol. 57, 1912, p. 175,
pl. 28, figs. 3, 4.--Cleland, Geology, Physical and Historical, New
York, 1916, p. 412, fig. 382 F (somewhat restored).
This very imperfectly known form is referred by Walcott to the
Notostraca on what appear to be wholly inadequate grounds, and while I
do not insist on my interpretation, I can not refrain from calling
attention to the fact that it _can_ be explained as the most primitive
of all trilobites. It consists of two subequal shields, the anterior
of which shows slight, and the posterior considerable evidence of
segmentation. It has no eyes, no glabella, and no thorax, and is
directly comparable to a very young _Phalacroma bibullatum_ (see
Barrande 1852, pl. 49, figs. a, b). Walcott states that there is
nothing to show how many segments there are in the cephalic shield,
but that on one specimen fourteen were faintly indicated on the
abdominal covering. The appendages are imperfectly unknown, as no
specimen showing the ventral side has yet been described. The possible
presence of antennas and three other appendages belonging to the
cephalic shield is mentioned, and there are tips of fourteen legs
projecting from beneath the side of one specimen. As figured, some of
the appendages have the form of exopodites, others of endopodites,
indicating that they were biramous.
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