The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
derived primitive Malacostraca retaining biramous antennæ, but with a
carapace and reduced pleural lobes and pygidium. From this offshoot
were probably derived the Ostracoda, the Cirripedia, and the various
orders of the Malacostraca, with the possible exception of the
Isopoda. I have suggested independent origins of the Copepoda and
Isopoda, but realize the weighty arguments which can be adduced
against such an interpretation.
[Illustration: Fig. 36.--_Naraoia compacta_ Walcott. An outline of
the test, after Walcott. Natural size.]
[Illustration: Fig. 37.--_Pagetia clytia_ Walcott. An eodiscid with
compound eyes. After Walcott. × 5.]
[Illustration: Fig. 38.--_Asaphiscus wheeleri_ Meek. A representative
trilobite of the Middle Cambrian of the Pacific province. After Meek.
× 1/2.]
[Illustration: Fig. 39.--_Pædeumias robsonensis_ Burling. Restored
from a photograph published by Burling. × 1/4.]
[Illustration: Fig. 40.--_Robergia_ sp. Restored from fragments found
in the Athens shale (Lower Middle Ordovician), at Saltville, Va.
Natural size.]
It is customary to speak of the Crustacea and Trilobita as having had
a common ancestry, rather than the former being in direct line of
descent from the latter, but when it can be shown that the higher
Crustacea are all derivable from the Trilobita, and that they possess
no characteristics which need have been inherited from any other
source than that group, it seems needless to postulate the evolution
of the same organs along two lines of development.
I can not go into the question of which are more primitive, sessile or
stalked eyes, but considering the various types found among the
trilobites, one can but feel that the stalked eyes are not the most
simple. While no trilobite had movable stalked eyes, it is possible to
homologize free cheeks with such structures. They always bear the
visual surface, and, in certain trilobites (_Cyclopyge_), the entire
cheek is broken up into lenses. Since a free cheek is a separate
entity, it is conceivable that it might lie modified into a movable
organ.
EVOLUTION OF THE MEROSTOMATA.
It has been pointed out above that the Limulava (_Sidneyia_,
_Amiella_, _Emeraldella_) have certain characteristics in common with
the trilobites on the one hand and the Eurypterida on the other. These
relationships have been emphasized by Walcott, who derives the
Eurypterida through the Limulava and the Aglaspina from the Trilobita.
The Limulava may be derived from the Trilobita, but indicate a line
somewhat different from that of the remainder of the Crustacea. In
this line the second cephalic appendages do not become antennæ and
the axial lobe seems to broaden out, so that the pleural lobes become
an integral part of the body. As in the modern Crustacea, the pygidium
is reduced to the anal plate, and this grows out into a spine-like
telson.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account