The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
In the Bohemian _Trinucleoides reussi_ (Barrande), the oldest of the
trinucleids, the hypostoma is very commonly present, and is of the
proper size to just cover the cavity of the glabella, seen from the
lower side, and has, toward the anterior end, side flaps which reach
out under the prominent glabellar lobes. This large size of the
hypostoma would cause the antennules to be attached outside the dorsal
furrows, and the position in which they are attached in the American
species of _Cryptolithus_ may be explained as an inherited one, since
with the small hypostoma they might have been within the glabella, as
in _Triarthrus_.
The antennules are seen in three specimens, and in all cases are
directed backward. The particular course in which they are drawn in
the restoration is purely arbitrary. The second pair of cephalic
appendages are represented as directed downward and forward, since in
one or two specimens fragments of forward-pointing endopodites were
seen near the front of the cephalon, and because in other trilobites
the second pair of appendages is always directed forward. The
remaining three pairs have a more solid basis in observed fact, for
the two or three specimens retaining fragmentary remains of them
indicate that they turn backward like those on the thorax, and that
the individual segments are longer and more nearly parallel-sided than
those of the more posterior appendages. The gnathites of all the
cephalic appendages are admittedly purely hypothetical. None of the
specimens shows them. As drawn, they are singularly inefficient as
jaws, but if, as is suggested by the casts of the intestines of
trinucleids found in Bohemia, these trilobites were mud-feeders,
inefficient mouth-parts would be quite in order.
[Illustration: Fig. 20. _Cryptolithus tessellatus_ Green. A
restoration of the appendages drawn by Doctor Elvira Wood from the
original specimens and from the photographs made by Professor Beecher.
× 9.]
The appendages of the thorax and pygidium can fortunately be taken
quite directly from the photographs of the dorsal and ventral sides of
well preserved specimens. There is of course a question as to the
number and the exact form of those on the pygidium, but I think the
present restoration is fairly well justified by the specimens. As
would be expected from the narrow axial lobe, the gnathobases of the
coxopodites are short and small.
Summary on the Ventral Anatomy of Trilobites.
COMPARISON OF APPENDAGES OF DIFFERENT GENERA.
Since the appendages of _Triarthrus_, _Cryptolithus_, _Neolenus_,
_Calymene_, and _Ceraurus_ are now known with some degree of
completeness, those of _Isotelus_ somewhat less fully, and something
at least of those of _Ptychoparia_, _Kootenia_, and _Acidaspis_, these
forms being representatives of all three orders and of seven different
families of trilobites, it is of some interest to compare the
homologous organs of each.
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