_Evolutionary history._--The genus _Megapodius_ consists, according to
Peters (1934:1-7), of nine species which are distributed through the
islands from the Philippines and Borneo to Australia and Melanesia.
These have been redesignated under three specific names by Mayr (1938).
Outlying forms occur in the Nicobar Group to the west and in Tonga
(Niuafou Island) in the east and in the Palaus and Marianas to the
extreme northeast. Lister (1911:757) is of the opinion that the
megapodes may have reached these outlying islands by having been
transported by the natives, by whom the eggs were highly valued as food.
This idea is also maintained by Rutland (1896:29-30) and Christian
(1926:260). Possibility and not factual evidence support this
hypothesis. From their seeming ancestral stocks, _M. pritchardii_ Gray
of Niuafou Island and _M. lapérouse_ of Micronesia are remarkably
distinct which may indicate their early arrival at these islands and
subsequent change from their ancestral stocks.
Like _M. pritchardii_, the Micronesian species is smaller than its
relatives to the southwest and has short, rather rounded wings, although
its feet are heavily built whereas those of _M. pritchardii_ are lightly
constructed. In comparing these birds with the species of megapode found
in the Philippines, Celebes and Melanesia, it seems that both _M.
pritchardii_ and _M. Lapérouse_ are closely related to the widespread
species, _M. freycinet_, which may have been ancestral to both. The
differences between _M. prichardii_ and _M. lapérouse_ indicate that
they represent independent invasions. Nevertheless these megapodes may
have had a wider range in Oceania in former times; man may have
eliminated the birds from some islands by using their eggs. The eggs are
laid in conspicuous mounds which are easily found by man.
_M. lapérouse_ differs from _M. freycinet_ of New Guinea and other parts
of Melanesia and the Philippines; its small size, short wing and pearl
gray head are distinctive characters. It shows greatest resemblance to
the subspecies in Celebes (_M. f. gilberti_) in size and to the
subspecies in the Moluccas (_M. f. freycinet_) in coloring; possibly
_M._ _lapérouse_ represents stock from one of these regions. Apparently
the group as a whole evolved from a center of dispersal in the New
Guinea area. Mayr (1942b:167) regarded all the species of _Megapodius_
as belonging to one polytypic species, except _M. lapérouse_ and _M.
pritchardii_, which are allopatric species.
=Coturnix chinensis lineata= (Scopoli)
Painted Quail
_Oriolus lineatus_ Scopoli, Del. Flor. et Faun. Insubr., fasc. 2,
1786, p. 87. (Type locality, Luzon, _ex_ Sonnerat.)
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