_Gygis a. microryhncha_ was for a long time treated as a species
distinct from _G. alba_ but has recently been considered as a subspecies
_G. alba_ by Peters and others. On the islands of Hatutu and Motane in
the Marquesas, the Whitney South Sea Expedition obtained some birds
which appear to be intergrades between the two subspecies of White
Terns in the area. The measurements of nine birds which show
intergradation between _G. a. microrhyncha_ and _G. a. pacifica_ are
listed in table 24. Probably the Marquesas population is tending toward
complete reproductive isolation.
[Illustration: FIG. 12. Geographic distribution of _Gygis alba_ in the
Pacific area. (1) _G. a. candida_; (2) _G. a. pacifica;_ (3) _G. a.
microrhyncha;_ (4) _G. a. royana_.]
Peters (1934:348, 349) recognizes three other subspecies from the
Pacific area: _G. a. rothschildi_ Hartert from Laysan, Lisiansky, and
Krusenstern islands; _G. a. candida_ (Gmelin) from "the Carolines east
to Christmas Island and south to the Tonga and Society Islands"; and _G.
a. royana_ Mathews from Norfolk and the Kermadec Islands. Birds from
Revilla Gigedo, Cocos and Clipperton islands, although geographically
isolated, are placed in _G. a. candida_. On the basis of a critical
study of specimens at hand, the populations in the Pacific fit into
three groups. Small birds, _G. a. candida_, are found in the North
Pacific from the Bonins and Marianas east to Wake and the Hawaiian Chain
and south to the Marshall, Phoenix, Christmas and the Fanning islands
(see figure 12). Larger birds, _G. a. pacifica_, are found in the
Central Pacific and South Pacific from the Carolines in the west
southeastward through Melanesia and eastward through Samoa, to the
Tuamotus and Easter to Cocos, Clipperton, and Revilla Gigedo islands. In
the Southwest Pacific, at Norfolk and the Kermadec Islands, a
longer-winged population occurs; it is separable as _G. a. royana_. The
measurements of these birds are given in table 24.
TABLE 24. MEASUREMENTS OF _Gygis alba_ FROM THE PACIFIC AREA
Column headings:
A: No.
B: Wing
C: Longest tail feather
D: Shortest tail feather
E: Exposed culmen
F: Depth Culmen
G: Tarsus
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