Palau owes its marked divergence to its isolated position at the
periphery of the range of the species. _P. p. pelewensis_ probably
represents an independent and an earlier colonization, possibly from a
stock different from that from which the two subspecies in the Carolines
arose. The presence in the Palaus of subspecies singularly different
from subspecies in the Carolines can also be observed in other genera,
as for example, _Rhipidura_, and _Myiagra_. Figure 13 shows the inferred
routes of colonization of _Ptilinopus_ to Micronesia.
_P. roseicapillus_ seemingly represents a remnant, or perhaps a
successful straggler, of an early invasion to Micronesia. Ripley and
Birckhead (1942:2) classify this species as "Old Stock," along with _P.
monachus_, _P. coronulatus_ and _P. regina_. Its pathway of invasion to
the Marianas was probably directly northward from the Papuan area and
not by way of the Polynesian islands. Its resemblance to the species _P.
regina_ of southern Papua, Lesser Sundas, and Australia is most unusual,
especially since there is a separation between the two species of some
1,400 miles; this is pointed out by Ripley and Birckhead (1942:4). As I
have said (1948:59) elsewhere, "On the basis of its characters the
Mariana birds would merit only subspecific separation, but owing to the
great distance between the two doves and the possibility of independent
origin and subsequent convergence, it may be more advisable to continue
to regard the two as separate species."
[Illustration: FIG. 13. Geographic distribution of _Ptilinopus
porphyraceus_ and routes of its dispersal. (1) _P. p. porphyraceus_; (2)
_P. p. fasciatus_; (3) _P. p. hernsheimi_; (4) _P. p. ponapensis_; (5)
_P. p. pelewensis_.]
=Ducula oceanica monacha= (Momiyama)
Micronesian Pigeon
_Globicera oceanica monacha_ Momiyama, Birds Micronesia, March,
1922, p. 4. (Type locality, Yap.)
_Columba oceanica_ Lesson and Garnot (part), Dict. Sci. Nat., éd.
Levrault, 40, 1826, p. 317 (Pelew); Lesson (part), Man. d'Ornith.,
2, 1828, p. 166 (Pelew); _idem_ (part), Voy. "La Coquille," Zool.,
2, 1828, pp. 432, 709 (Pelew); _idem_, Compl. de Buffon, 2d ed.,
2, Oiseaux, 1838, p. 292 (Pelew); Prévost and Knip, Les Pigeons,
2, 1838-43, p. 49 (Pelew).
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