_Evolutionary history of Corvus kubaryi._--Meinertzhagen (1926:59)
writes that "Environmental influences seem to be mainly, if not
entirely, responsible for geographic differences in the genus _Corvus_."
Such may be the case in _C. kubaryi_, which is a small, dull-colored
crow with a relatively unmodified bill. In structure, it has little
resemblance to other crows found in the Pacific area. Kittlitz was the
first to note a resemblance between the bird at Guam and one in the
Philippines. Oustalet (1896:70) wrote that the bird at Guam is related
to crows of the Moluccas and New Guinea. Although not closely related to
the Hawaiian Crow, _C. tropicus_, both have little gloss on their
feathers, a character which is common to many of the insular populations
of crows. Mayr (1943:46) is of the opinion that the Hawaiian bird was
derived from a North American ancestor, although Bryan (1941:187)
suggests that it is related to _C. macrorhynchus_ of southeastern Asia
and remarks that the Hawaiian Crow, "has some relation to the Guam
Crow." In looking for the ancestral stock of _C. kubaryi_, the several
species of crows which occur to the north, west and south of the
Marianas have been examined. In size and general structure, _C. kubaryi_
appears to be closest to the _C. enca_ group, and not as closely related
to the _C. macrorhynchus_ group. The small size, the shape of the
culmen, the lack of pointed feathers on the breast, and the presence of
white on the basal parts of the feathers of the nape are characters
which _C. kubaryi_ has in common with _C. enca_. Nasal bristles cover
the frontal base of the culmen in _C. kubaryi_; this character is found
also in _C. enca florensis_. _C. kubaryi_ differs from the _C. enca_
group by lacking the purple sheen on the upper parts; this sheen is
conspicuous in the latter species. _C. kubaryi_ appears to have little
in common with _C. meeki_ of the Solomons and _C. orru_ of the Moluccas
and New Guinea area. There is apparently no close relation between the
Marianas Crow and the crow which reaches the Bonins. The latter crow,
according to the Hand-list of Japanese Birds (Hachisuka _et al._,
1932:1), is called _C. coronoides hondoensis_ Momiyama and is apparently
now extinct in the Bonins.
In summary, it may be said that _C. kubaryi_ is an isolated and modified
species of crow, which probably has been living at Guam and Rota for a
considerable length of time. Whether it once lived on other islands in
Micronesia is unknown, but it is entirely possible that the present
population may represent a remnant of one which formerly had a more
extensive distribution. The characters which show its distinctness from
possible ancestral species include its small size, its slender bill,
and its dull coloration. It is thought to have been derived from the _C.
enca_ group, _C. e. pusillus_ of the Philippines or _C. e. celebensis_
of the Celebean area.
=Luscinia calliope calliope= (Pallas)
Siberian Rubythroat
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