The Biological Problem of To-day: Preformation Or Epigenesis?: The Basis of a Theory of Organic DevelopmentHertwig, Oscar
Science
The Biological Problem of To-day: Preformation Or Epigenesis?: The Basis of a Theory of Organic Development
Hertwig, Oscar
Developmental biology; Embryology; Genetics
Thirdly, great stress is to be laid on those experiments in which the
process of development was interfered with at different stages, as these
showed that the separate cells which arose by division were not predestined
unalterably for a particular _rôle_, according to a predetermined plan
(facts of regeneration and heteromorphosis).
Fourthly, the results of grafting, transplantation, and transfusion
indicate that the cells and tissues of an organism possess, in addition to
their patent microscopical characters, latent characters, which show
themselves to be peculiar to the species.
How does Weismann attempt to reconcile his hypothesis of differentiating
division with these facts? By the provision of different complementary
hypotheses, which, as we have seen, amount to this, that he allows the set
of rudiments which he had turned out by differentiating division of the
cell to creep in again by a back-door. He accomplishes this by his idea
that the germplasm may undergo, simultaneously, doubling and
differentiating division. In these cases cell-division has a double aspect.
According to Weismann, this is possible, because the egg contains many,
sometimes as many as a hundred, _ids_, each of which is a combination
representing the species. Weismann believes that in an egg, while it is
preparing for its first division, the _ids_ are arranged in two groups--an
active army and a reserve army. By differentiating division the active army
is broken up into the divisions, brigades, and regiments of determinants
appropriate to the separate groups of cells, and so the course of the
development is conducted according to a preconceived plan. On the other
hand, the passive, reserve army multiplies by doubling division, and is
sent along with definite parts of the active army as baggage in a fixed or
inactive condition, so that it has no influence upon the normal course of
development nor upon the characters of the cells (fixed germplasm,
inactive, accessory idioplasm, bud-idioplasm).
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account