The Biological Problem of To-day: Preformation Or Epigenesis?: The Basis of a Theory of Organic DevelopmentHertwig, Oscar
Science
The Biological Problem of To-day: Preformation Or Epigenesis?: The Basis of a Theory of Organic Development
Hertwig, Oscar
Developmental biology; Embryology; Genetics
As I see Nature, this contrast has been artificially reasoned into her.
From several reasons, I do not think that it exists. In the first place, I
consider that the facts I have given show the hypothesis of a
differentiating division of cells and germplasm to be not proven and
arbitrary. Next, the reproductive-cells must be considered as much a part
of the organism as any other tissue. Sometimes they form the greater part
of the body, as in many parasites, and, like the other tissues, they are
subject to death, unless the conditions necessary to their further
development have occurred in time. But under such conditions other
cell-complexes may have death averted from them, as, for instance, when a
slip cut from a willow-tree is planted. Thirdly, the reproductive cells are
derived from the egg-cell just in the same way as other tissue cells are
derived from it. Like tissue cells in multicellular organisms, they arise
by the specialisation of material separated from the egg-cell, and, like
every other organ, attain the position assigned them in the plan of
development in the course of the general metamorphosis of position that all
the cells pass through. Often the sexual cells, like those of other
tissues, appear at a distance of several cell-generations from the egg. The
intervening generations are specially numerous in those animals and plants
in which several sexless generations come between the sexual generations
(_e.g._, many plants, coelenterates, worms, tunicates).
I cannot agree to the existence (in Weismann's sense) of special
germ-tracks. Naturally, I do not deny that the sexual cells arise from the
egg after definite sequences of cell-divisions; but this happens in the
case of all specialised cells, such as muscle, liver, kidney, and bone
cells. The conception of special germ-tracks has no more significance than
there would be in the conception of muscle, liver, kidney, and bone
tracks. Though Weismann associates with germ-tracks the idea that germplasm
travels along them, proof of this has yet to be brought forward.
Finally, a word about the meaning of 'immortal.' In a scientific work the
word must be used in a philosophical sense. In calling a being immortal one
implies both individuality and indivisibility. This, at least, was the view
of the old philosophers, who have defined the idea of immortality. Thus
says Leibnitz in his _Theodice_: 'I hold that the souls which one day
become the souls of men existed already in the seed, that they have existed
always in organised form in the ancestors, back to Adam--that is to say, to
the beginning of things.'
Public-domain text, read in full here on John Shaqi.
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