The Biological Problem of To-day: Preformation Or Epigenesis?: The Basis of a Theory of Organic DevelopmentHertwig, Oscar
Science
The Biological Problem of To-day: Preformation Or Epigenesis?: The Basis of a Theory of Organic Development
Hertwig, Oscar
Developmental biology; Embryology; Genetics
bringing together the different materials absorbed by roots and leaves; and
they have the mechanical function of strengthening the stem and branches.
The different mode of nutrition of animals results in a totally different
structural plan. Animal cells absorb material that is already organised,
and that they may do so their cells are either quite naked, so affording an
easy passage for solid particles, or they are clothed only by a thin
membrane, through which solutions of slightly diffusible, organic colloids
may pass. Therefore, unlike plants, multicellular animals display a compact
structure with internal organs adapted to the different conditions which
result from the method of nutrition peculiar to animals. A unicellular
animal takes organic particles bodily into its protoplasm, and forming
around them temporary cavities known as food vacuoles, treats them
chemically. The multicellular animal has become shaped so as to enclose a
space within its body into which solid organic food-particles are carried
and digested, thereafter, in a state of solution, to be shared by the
single cells lining the cavity. In this way the animal body does not
require so close a relation with the medium surrounding it; its food, the
first requirement of an organism, is distributed to it from inside
outwards. In its further complication the animal organisation proceeds
along the same lines. The system of internal hollows becomes more
complicated by the specialisation of secreting surfaces, and by the
formation of an alimentary canal, and of a body cavity separate from the
alimentary canal.
In plants, it is the external surface that is increased as much as
possible. In animals, in obedience to their different requirements,
increase takes place in the internal surface. The specialisation of plants
displays itself in organs externally visible--in leaves, twigs, flowers,
and tendrils. The specialisation of animals is concealed within the body,
for the internal surface is the starting-point for the formation of the
organs and tissues.
Comparative embryology shows that, however varied the forms and functions
of the numerous animal organs may be, the method of their development is
remarkably similar. There are required only the slightest variations of a
few simple general laws. For these I may refer readers to a series of
special investigations (_Studies on the Germ-layer Theory_, Oscar and
Richard Hertwig), and to the fourth chapter of my Embryology, 'General
Discussion of the Principles of Development.'
Public-domain text, read in full here on John Shaqi.
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