The Body at Work: A Treatise on the Principles of PhysiologyHill, Alex
Science
The Body at Work: A Treatise on the Principles of Physiology
Hill, Alex
Physiology
My lord the stomach! He is not the only, nor is he the chief, agent
in digestion; but with him rests the decision as to whether the food
offered to the alimentary tract is suitable in quality and quantity. He
is offended if it be not offered with all the circumstance and ceremony
which becomes his rank. As an intimation that he is about to receive
food, he accepts the news from the mouth that its nerve-endings are
subject to mechanical stimulation. But the chewing of indiarubber would
produce a like effect. The stomach, therefore, confers with the organs
of taste and smell. If their report is favourable, he argues that the
substance which the teeth are crushing will justify an outflow of
gastric juice. He responds most generously when prolonged mastication
assures him that he may trust to receiving the food in a sufficiently
subdivided state. At our peril we neglect to propitiate my lord. Not
always debonair when treated with consideration, he is morose or
petulant when slighted. Never content with lip-service, he exacts the
labour of teeth and tongue and palate. The tribute we offer may be of
the best—savoury, wholesome, well cooked, well chewed—but if it be
not tendered with some degree of love, if thoughts are concentrated
on other things, if no attention is devoted to the meal, if no sense
of liking accompanies our offering, my lord the stomach on his part
affords the viands an indifferent reception. In consulting our own
tastes we are to a large extent consulting the needs of the stomach.
Ravenous and excessive feeding is not an exhibition of taste; it is a
return to the instinct of the savage, who was never sure that he would
get his full share, and was afraid to trust that another meal would be
obtainable when nature declared it due. Some degree of epicureanism
is favourable to digestion. The flow of gastric juice in the stomach
occurs reflexly in response to the emotion of appetite, to stimulation
of the nerves of taste and smell, to the obscure sensations which
accompany the activity of the muscles of mastication.
The gastric juice secreted in a day amounts probably to about 8 or 9
pints. To this we must add, when considering the quantity of fluid
which passes through the stomach, the saliva, which certainly reaches
as much as 2 pints, and the beverages taken with food.
Gastric juice collected in the manner described above is a clear,
colourless, inodorous fluid. It is very acid, and so powerfully peptic
as to digest its own weight of coagulated white of egg. Its solid
constituents amount to 0·5 per cent. They consist of the two ferments
pepsin and rennin, with traces of proteins and mucin, and various
inorganic salts. Its acidity is due to free hydrochloric acid to the
amount of 0·2 per cent. This acid is more or less in combination with
the pepsin. In pure gastric juice hydrochloric acid is the only acid
present; but when mixed with food the juice contains other acids also,
especially lactic.
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