The Body at Work: A Treatise on the Principles of PhysiologyHill, Alex
Science
The Body at Work: A Treatise on the Principles of Physiology
Hill, Alex
Physiology
The valve “balloons” over the blood in the ventricle before the
contraction of the auricle has ceased. The thin margins of its flaps
come together with great rapidity. The tendinous cords holding their
edges on the ventricular side, they meet, not edge to edge, but folded
flap to folded flap. (3) The valve does not bulge into the auricle. On
the contrary, at the height of systole it is pulled into the ventricle
by the contracting musculi papillares. As the ring to which the valve
is attached is diminished in size, by the contraction of the base of
the heart, which continues, it will be remembered, until after the apex
has begun to relax, the edges of the flaps are folded farther and still
farther over by the pull of the musculi papillares, and the blood is
squeezed out from between the wall of the ventricle and the indrawn
valve.
The “semilunar valves,” which close the apertures into the aorta and
pulmonary artery, have each three flaps. The aortic semilunar valve,
which has the higher pressure to bear, shows its characteristic
features in a rather more marked degree than the other. Each of its
three flaps is a half-cup. At the centre of the margin of the half-cup
is a small fibrous nodule. The edge of the cup on either side of this
is very thin. Fine elastic fibres radiate from the nodule to all parts
of the flap. The wall of the aorta shows three bays, or “sinuses,”
one behind each flap. Hence, when the valve is forced by the rise of
pressure in the ventricle, the flap is not flattened back against the
wall of the aorta. There is always a certain amount of backwash in the
pocket behind it. The instant the pressure in the ventricle begins
to fall, the three flaps come together with a click, so smart as to
be plainly audible over most of the front of the chest. The click is
the “second sound” of the heart. The auriculo-ventricular valves also
make a sound when they close; but this “first sound of the heart”
has a different character. It is prolonged, soft, low-pitched. It is
customary to represent the sounds by the syllables “lūbb dŭp—lūbb
dŭp,” the pause during diastole being of about the same length as
the sounds when the heart is beating with its normal rhythm. The
duration of systole is little affected by variations in the rate of
beat. It is diastole that is shortened or prolonged. The second sound
is due entirely to the closure of the semilunar valves. It is heard
most clearly when the stethoscope is placed over the region where
the aorta comes nearest to the wall of the chest—at the second rib
cartilage on the right side of the breast-bone. The first sound is
loudest near the apex of the heart. It is generally agreed that it is
not wholly due to the closure of the auriculo-ventricular valves, but
possesses a second constituent. Some persons assert that they can with
the ear distinguish the clearer valvular sound at the commencement
from the general rumble which overtakes it. The main part of the
Public-domain text, read in full here on John Shaqi.
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