The Body at Work: A Treatise on the Principles of PhysiologyHill, Alex
Science
The Body at Work: A Treatise on the Principles of Physiology
Hill, Alex
Physiology
The third variety of muscle is the kind by which locomotion is
effected. It is present in large masses—all the red tissue to which
the term “meat” is commonly applied. It accounts for about 35 per cent.
of the body-weight. This kind of muscle is not composed of single
cells, but of compound cells, or cell-complexes, termed “fibres.” A
fibre may attain a length of upwards of 2 inches, with a breadth of
about ¹/₅₀₀ inch. In most cases the fibres are attached by one end to
a bone, by the other to a tendon; and since they are shorter than the
muscle as a whole, the tendon commences as a membrane which covers the
surface of the muscle, sloping to it from the bone to which by their
other ends the fibres are attached. A fibre is developed from a single
cell. The cell elongates, its nucleus divides, and the daughter-nuclei
divide until several hundred have been formed; but cell division
does not follow. The result is a cylindrical mass enclosed within a
delicate membranous sheath, the sarcolemma. In the early stages of its
development its nuclei are in the axis of the fibre, but subsequently
they are displaced outwards. In the most highly specialized muscle,
known as the “white” variety, they lie just beneath the sarcolemma
(_cf._ Fig. 16, B).
The feature of this type of muscle is its transverse striation, almost
mathematically regular. Commonly striated muscle is spoken of as
“voluntary,” because, for the most part, it is under the control of
the Will; but the term, in so far as it implies a connection between
structure and mode of actuation, is misleading. Transverse striation
is evidence of capacity for rapid action. The muscles which the Will
directs exhibit promptitude; but striated muscle, which is not under
the direction of the Will, is found in certain situations—_e.g._, the
upper part of the œsophagus. Conversely, many animals can voluntarily
call into action muscle which is not striped. A turkey erects its
feathers by setting in motion little groups of “plain” fibres, which
pull on elastic tendons attached to the tips of the buried ends of
their shafts. Plain muscle contracts less promptly and relaxes more
slowly than the striped variety. Cardiac muscle is quicker in acting
than plain, but does not hold the contraction so long.
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