The Body at Work: A Treatise on the Principles of PhysiologyHill, Alex
Science
The Body at Work: A Treatise on the Principles of Physiology
Hill, Alex
Physiology
=Elasticity of Muscles.=—Muscles are very extensible, and after
stretching return to their original length. Their elasticity is a
quality of great practical importance. It enables them to meet sudden
resistance without rupture, as when a man alights from a height. At
the moment when the feet touch ground elasticity dissipates the shock.
The stretching of the muscles then leads reflexly to the increase of
their tone. Here we see an advantage in the short reaction-time of
the knee-jerk. Tone comes into play long before impulses generated by
contact of the sole of the foot with the ground have had time to reach
the brain, or even to induce reflex contraction through the spinal
cord. The elasticity of the muscles is also of use in the performance
of certain sudden actions. A pea is flicked across the room by pressing
the thumbnail against the pad of a finger, or a finger against the
thumb, and releasing it with a jerk.
An electrical change accompanies an impulse in its passage down a
nerve, and a wave of contraction in its passage along a muscle. In
1788 Galvani observed that the hind-limbs of a frog, suspended by
a metal hook to metal railings, twitched when the wind blew them
against the bars. The hook passed through the lumbar plexus of
nerves. He recognized that the cause of the twitch was the closing of
a circuit. The birth of dynamic or galvanic electricity dates from
this observation; and ever since this phenomenon was first observed
the electric changes in nerve-muscle preparations made from frogs’
legs have been favourite subjects of research. Many observations with
regard to nerve-conduction and muscle-contraction may be made, and many
experiments performed, without special apparatus. A frog having been
killed by cutting off its head, or by placing it beneath a tumbler
with a wad of cotton-wool soaked in chloroform, the skin of the leg
is removed, displaying the khaki-coloured muscles, bluish tendons,
and bright white threads of nerve. A stretch of the largest nerve of
the back of the thigh, the sciatic, is isolated. All the muscles of
the thigh are then cut away and the bone nipped across just above the
knee. The bones below the knee are removed, the superficial muscle of
the calf, the gastrocnemius, being allowed to hang free, its bifid end
attached to the fragment of thigh-bone. Its lower end terminating in
the tendo Achillis, with its insertion into the prominence of the heel,
is left intact. The bone is fixed in a clamp. A light lever made from
a wooden spill is suspended from the tendo Achillis. The nerve may
then be stimulated in various ways: by crushing in a pair of forceps,
burning with a heated needle, touching with a drop of glycerin or
a strong solution of salt. But of all methods of stimulation, the
best is the current from an induction coil. Since it does not injure
the nerve, it can be applied as often as may be desired. The amateur
provided with an induction coil is in a position to study the relation
Public-domain text, read in full here on John Shaqi.
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