The Body at Work: A Treatise on the Principles of PhysiologyHill, Alex
Science
The Body at Work: A Treatise on the Principles of Physiology
Hill, Alex
Physiology
Usually the curve shows a rise lasting ⁴/₁₀₀ second and
a fall occupying ⁵/₁₀₀, due to the fact that inertia of muscle and
apparatus delays the commencement of the rise, but does not hasten the
termination of the fall.
When an impulse is generated artificially by an induction shock, a
single spasm or twitch is the result; but in Nature contraction is
never limited to a single twitch. Impulses descending from a motor
nerve-cell to a muscle are always rhythmic. They follow at the rate
of eight or ten a second in human nerves; and since in our muscles
contraction and relaxation take longer than in a frog, a second impulse
reaches the muscle before the effect of the first has passed away.
The muscle has not had time to relax, when it is again called upon to
contract. Hence a summation of contractions. The muscle continues to
shorten until the maximum of contraction is reached. This condition is
termed “tetanus,” to distinguish it from a single spasm. In fullest
contraction the length of a muscle may be diminished by one-half, or
even by two-thirds.
It would be impossible to treat of the =electrical phenomena= displayed
by nerves and muscles without presupposing some acquaintance with the
methods and laws of physics. As this is contrary to our understanding
with our readers, we must be content with the statement of a few
salient facts. At the moment when an impulse is passing along a nerve,
or a wave of contraction along a muscle, the electric potential of the
active part of the structure, whether nerve or muscle, is different
from that of the not-acting parts on either side of it. A battery in
its commonest form is a glass vessel containing sulphuric acid in
which a plate of zinc and a plate of copper are immersed. The zinc is
electro-positive as regards the copper. In a muscle the contracted
portion is electro-positive as regards the parts uncontracted. The
degree of positivity can be measured by connecting the muscle at two
spots with the two wires of a galvanometer. When one wire makes contact
with the contracted portion, and the other with a part which is not
contracted, a current passes through the galvanometer, causing its
needle to swing; and since the wave of contraction is not stationary,
but passes down the muscle, the current is subsequently reversed.
The wave, as it were, first tilts up one end, and then, passing on,
tilts up the other, letting down the first. The contracted spot is
electro-positive to the spot not contracted, and then the latter,
contracting, becomes electro-positive to the former, which has relaxed.
The needle of the galvanometer swings first to the left, then to the
right. The importance of this method of investigation lies in the fact
that the electric variation exactly represents, both in time and in
intensity, the change which is occurring in nerve and in muscle. By
following it, we can ascertain the rate at which an impulse travels
down a nerve. We can determine its length and its “form.” Represented
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