The Body at Work: A Treatise on the Principles of PhysiologyHill, Alex
Science
The Body at Work: A Treatise on the Principles of Physiology
Hill, Alex
Physiology
He is hardly prepared to say that its
greater vivacity may not be the expression of more rapid oxidation;
but he sees no fore-ordained balance of vital enterprise and chemical
change. He is, or ought to be, extremely suspicious of any explanation
which appears to over-ride physical laws; yet, at the same time, he
is aware that until he has more accurate knowledge regarding the
constitution of protoplasm he will not be in a position to understand
how physical laws apply. The protoplasmicity of protoplasm is increased
by warmth. What change of molecular constitution does this imply?
The view that in a muscle molecular change gives rise to an electrical
change, which in turn produces the change in form, has been very
widely held. The hypothesis was based on observations which seemed
to show that the electric variation travels a little ahead of the
wave of contraction; but every improvement in recording apparatus has
diminished this apparent want of synchronism. There can be little doubt
but that the lagging behind of the wave of contraction is due to the
inertia of the muscle and of the recording apparatus. Molecular change
and electric variation are simultaneous. If this be true, the electric
change cannot be regarded as the cause of the molecular change, in the
sense, at any rate, in which they used to be considered as cause and
effect.
The =power of muscle= varies as its cross-section. For human muscles
the maximum lift amounts to from 7 to 10 kilogrammes for each square
centimetre. This is a large figure, but it must be remembered that,
owing to the arrangement of the bones as levers, most muscles act
at a great mechanical disadvantage. The greater the difference in
distance from the fulcrum between the point of application of the
force and the point of incidence of the weight, when the force acts
nearer to the fulcrum than the weight, the greater is the mechanical
disadvantage. The greater also is the rapidity with which the weight
is lifted. What is lost in strength is gained in swiftness. Contrast
the slow steps of a negro, whose long heel separates the point of
application of the power (tendo Achillis) from the fulcrum (the
ankle-joint), with the springy movements of a European. A European
needs, and as a rule has, a better developed calf, which allows him
his more sprightly gait, without sacrificing his carrying power. Our
preference for slender wrists and ankles is not purely æsthetic, unless
we admit, as may be maintained, that all natural canons of taste rest
upon utility. Slimness of joints means nimbleness. A few muscles act
directly, without loss of power—as, for example, the masseter, which
lifts the lower jaw (hence a grand capacity for cracking nuts)—but
most muscles move levers of considerable length. Compare with the
masseter the biceps and brachialis which lift the forearm. Their
tendons are inserted into the radius and the ulna at a distance from
the elbow-joint which is about one-tenth as great as the distance from
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