The Botanic Garden, a Poem in Two Parts. Part 1: the Economy of VegetationDarwin, Erasmus
History
The Botanic Garden, a Poem in Two Parts. Part 1: the Economy of Vegetation
Darwin, Erasmus
Natural history; Plants -- Poetry; Portland Vase
At the same time so minute are the vessels in the intertexture of the
barks of plants, which belong to each individual bud, that a general
circulation may possibly exist, though we have not yet been able to
discover the venous part of it.
There is however another part of the circulation of vegetable juices
visible to the naked eye, and that is in the corol or petals of flowers,
in which a part of the blood of the plant is exposed to the influence of
the air and light in the same manner as in the foliage, as will be
mentioned more at large in Notes XXXVII and XXXIX.
These circulations of their respective fluids seem to be carried on in
the vessels of plants precisely as in animal bodies by their
irritability to the stimulus of their adapted fluids, and not by any
mechanical or chemical attraction, for their absorbent vessels propel
the juice upwards, which they drink up from the earth, with great
violence; I suppose with much greater than is exerted by the lacteals of
animals, probably owing to the greater minuteness of these vessels in
vegetables and the greater rigidity of their coats. Dr. Hales in the
spring season cut off a vine near the ground, and by fixing tubes on the
remaining stump of it, found the sap to rise twenty-one feet in the tube
by the propulsive power of these absorbents of the roots of it. Veget.
Stat. p. 102. Such a power can not be produced by capillary attraction,
as that could only raise a fluid nearly to the upper edge of the
attracting cylinder, but not enable it to flow over that edge, and much
less to rise 21 feet above it. What then can this power be owing to?
Doubtless to the living activity of the absorbent vessels, and to their
increased vivacity from the influence of the warmth of the spring
succeeding the winter's cold, and their thence greater susceptibility to
irritation from the juices which they absorb, resembling in all
circumstances the action of the living vessels of animals.
NOTE XXXVII--VEGETABLE RESPIRATION.
_While spread in air the leaves respiring play._
CANTO IV. l. 421.
I. There have been various opinions concerning the use of the leaves of
plants in the vegetable oeconomy. Some have contended that they are
perspiratory organs; this does not seem probable from an experiment of
Dr. Hales, Veg. Stat. p. 30. He found by cutting off branches of trees
with apples on them, and taking off the leaves, that an apple exhaled
about as much as two leaves, the surfaces of which were nearly equal to
the apple; whence it would appear that apples have as good a claim to be
termed perspiratory organs as leaves. Others have believed them
excretory organs of excrementious juices; but as the vapour exhaled from
vegetables has no taste, this idea is no more probable than the other;
add to this that in moist weather, they do not appear to perspire or
exhale at all.
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