The Botanic Garden, a Poem in Two Parts. Part 1: the Economy of VegetationDarwin, Erasmus
History
The Botanic Garden, a Poem in Two Parts. Part 1: the Economy of Vegetation
Darwin, Erasmus
Natural history; Plants -- Poetry; Portland Vase
[_Golden bell_. l. 460. There are muscles placed about the footstalks of
the leaves or leaflets of many plants, for the purpose of closing their
upper surfaces together, or of bending them down so as to shoot off the
showers or dew-drops, as mentioned in the preceeding note. The claws of
the petals or of the divisions of the calyx of many flowers are
furnished in a similar manner with muscles, which are exerted to open or
close the corol and calyx of the flower as in tragopogon, anemone. This
action of opening and closing the leaves or flowers does not appear to
be produced simply by _irritation_ on the muscles themselves, but by the
connection of those muscles with a _sensitive_ sensorium or brain
existing in each individual bud or flower. 1st. Because many flowers
close from the defect of stimulus, not by the excess of it, as by
darkness, which is the absence of the stimulus of light; or by cold,
which is the absence of the stimulus of heat. Now the defect of heat, or
the absence of food, or of drink, affects our _sensations_, which had
been previously accustomed to a greater quantity of them; but a muscle
cannot be said to be stimulated into action by a defect of stimulus. 2.
Because the muscles around the footstalks of the subdivisions of the
leaves of the sensitive plant are exerted when any injury is offered to
the other extremity of the leaf, and some of the stamens of the flowers
of the class Syngenesia contract themselves when others are irritated.
See note on Chondrilla, Vol. II. of this work.
From this circumstance the contraction of the muscles of vegetables
seems to depend on a disagreeable _sensation_ in some distant part, and
not on the _irritation_ of the muscles themselves. Thus when a particle
of dust stimulates the ball of the eye, the eye-lids are instantly
closed, and when too much light pains the retina, the muscles of the
iris contract its aperture, and this not by any connection or consent of
the nerves of those parts, but as an effort to prevent or to remove a
disagreeable sensation, which evinces that vegetables are endued with
sensation, or that each bud has a common sensorium, and is furnished
with a brain or a central place where its nerves were connected.]
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