The Cambridge natural history, Vol. 01 (of 10)Hickson, Sydney J. (Sydney John)
Science
The Cambridge natural history, Vol. 01 (of 10)
Hickson, Sydney J. (Sydney John)
Animals
Among the Euglenaceae, _Euglena viridis_ is a very common form, giving the
green colour to stagnant or slow-flowing ditches and puddles in light
places, especially when contaminated by a fair amount of dung, as by the
overflow of a pig-sty, in company with a few hardy Rotifers, such as
_Hydatina senta_ (Vol. II. Fig. 106, p. 199) and _Brachionus_. _Euglena_ is
about 0.1 mm. in length when fully extended, oval, pointed behind,
obliquely truncate in front, with a flagellum arising from the pharyngeal
pit. It shows a peculiar wriggling motion, waves of transverse constriction
passing along the body from end to end, as well as flexures in different
meridians. Such motions are termed "euglenoid." The front part is
colourless, but under a low {125}power the rest of the cell is green, owing
to the numerous chlorophyll bodies or chloroplasts. The outermost layer of
the cytoplasm shows a somewhat spiral longitudinal striation, possibly due
to muscular fibrils. The interior contains many laminated plates of
paramylum, and a large single nucleus. At the front of the body at the base
of the flagellum is a red "eye-spot" on the dorsal side of the pharynx-tube
or pit, from which the flagellum protrudes. Wager has shown that this tube
receives, also on its dorsal side, the opening of a large vacuole,
sometimes called the reservoir, for into it discharges the contractile
vacuole (or vacuoles). The eye-spot is composed of numerous granules,
containing the vegetal colouring matter "haematochrome." It embraces the
lower or posterior side of the communication between the tube and the
reservoir. The flagellum has been traced by Wager through the tube into the
reservoir, branching into two roots where it enters the aperture of
communication, and these are inserted on the wall of the reservoir at the
side opposite the eye-spot. But on one of the roots near the bifurcation is
a dilatation which lies close against the eye-spot, so that it can receive
the light reaction. _Euglena_ is an extremely phototactic organism. It
shows various wrigglings along the longitudinal axis, and transverse waves
of contraction and expansion may pass from pole to pole.[138]
Among the Chrysomonadaceae the genus _Zooxanthella_, Brandt, has already
been described under the Radiolaria (p. 86), in the jelly of which it is
symbiotic. It also occurs in similar union in the marine Ciliates,
_Vorticella sertulariae_ and _Scyphidia scorpaenae_, and in _Millepora_ (p.
261) and many Anthozoa (pp. 373 f., 396).
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