The Cambridge natural history, Vol. 01 (of 10)Hickson, Sydney J. (Sydney John)
Science
The Cambridge natural history, Vol. 01 (of 10)
Hickson, Sydney J. (Sydney John)
Animals
The NUCLEAR APPARATUS lies at the inner boundary of the ectoplasm. The
"meganucleus" may be ovoid, elongated, or composed of two or more rounded
lobes connected by slender bridges (_Stentor_, _Stylonychia_). The
"micronucleus" may be single; but even when the meganucleus is not lobed it
may be accompanied by more than one micronucleus, and when it is lobed
there is at least one micronucleus to each of its lobes.[160] The
meganucleus often presents distinct granules of more deeply staining
material, varying with the state of nutrition; these are especially visible
in the band-like meganuclei of the Peritrichaceae (Figs. 51, 60). At the
approach of fission it is in many cases distinctly fibrillated.[161] But
all other internal differentiation, as well as any constriction, then
disappears; and the ovoid or rounded figure becomes elongated and
hour-glass shaped, and finally constricts into two ovoid
daughter-meganuclei, which, during and after the fission of the cell,
gradually assume the form characteristic of the species. The micronuclei
(each and all when they are multiple) divide by modification of
karyokinesis (or "mitosis") as a prelude to fission: in this process the
chromatin is resolved into threads which divide longitudinally, but the
nuclear wall {145}remains intact. If an Infusorian be divided into small
parts, only such as possess a micronucleus and a fragment of the
meganucleus are capable of survival. We shall see how important a part the
micronuclei play in conjugation, a process in which the old meganuclei are
completely disorganised and broken up and their débris expelled or
digested.
The MOUTH of the Gymnostomaceae is habitually closed, opening only for the
ingestion of the living Protista that form their prey. It usually opens
into a funnel-shaped PHARYNX, strengthened with a circle of firm
longitudinal bars, recalling the mouth of an eel-trap or lobster-pot
("Reusenapparat" of the Germans); and this is sometimes protrusible. In
_Dysteria_ the rods are replaced by a complicated arrangement of jaw- or
tooth-like thickenings, which are not yet adequately described. We have
above noted the strong adoral trichocysts in this group.
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