The ASTROSCLERIDAE, as known at present, contain a single genus and species, apparently the most isolated in the phylum. _Astrosclera willeyana_[228] was brought back from the Loyalty Islands, and from Funafuti of the Ellice group. Its skeleton is both chemically and structurally aberrant. In other Calcarea the calcium carbonate of the skeleton is present as calcite, in _Astrosclera_ as aragonite, and the elements are solid polyhedra, {195}united by their surfaces to the total exclusion of soft parts (Fig. 88). Each element consists of crystalline fibres radially disposed around a few central granules, and terminating peripherally in contact with the fibres of adjacent elements. Young polyhedra are to be found free in the soft parts at the surface. The chambers are exceptionally minute, especially for a calcareous sponge, comparing with those of other sponges as follows:— _Astrosclera_ chambers, 10µ × 8µ to 18µ × 11µ. Smallest chambers in Silicea, 15µ × 18µ to 24µ × 31µ. Smallest chambers in Calcarea, 60µ × 40µ. In its outward form _Astrosclera_ resembles certain Pharetronids. The minute dimensions of the ciliated chambers relegate _Astrosclera_ to the Micromastictora, and the fortunate fact that the calcium carbonate of its skeleton possesses the mineral characters not of calcite, but of aragonite, renders it less difficult to conceive that its relations may be rather with the non-calcareous than the calcareous sponges. BRANCH II. MICROMASTICTORA All sponges which do not possess calcareous skeletons are characterised by choanocytes, which, when compared with those of Calcarea, are conspicuous for their smaller size. The great majority (Silicispongiae) of the non-calcareous sponges either secrete siliceous skeletons or are connected with siliceous sponges by a nicely graded series of forms. The small remainder are entirely askeletal. All these non-calcareous sponges are included, under the title Micromastictora, in a natural group, opposed to the Megamastictora as of equal value. The subdivision of the Micromastictora is a matter of some difficulty. The Hexactinellida alone are a well circumscribed group. After their separation there remains, besides the askeletal genera, an assemblage of forms, the Demospongiae, which fall into two main tribes. These betray their relationship by series of intermediate types, but a clue is wanting which shall determine decisively the direction in which the series are to be read. The askeletal genera are the _crux_ of the systematist. It is perhaps safest, while recognising that many of them bear a likeness of {196}one kind or another to various Micromastictora, to retain them together in a temporary class, the Myxospongiae. CLASS I. MYXOSPONGIAE The class Myxospongiae is a purely artificial one, containing widely divergent forms, which possess a common negative character, namely, the absence of a skeleton. As a result of this absence they are all encrusting in habit.