The Cambridge natural history, Vol. 01 (of 10)Hickson, Sydney J. (Sydney John)
Science
The Cambridge natural history, Vol. 01 (of 10)
Hickson, Sydney J. (Sydney John)
Animals
[Illustration: FIG. 93.—Scheme to show the arrangement of spicules in the
Hexactinellid skeleton. _Canalaria_, microscleres in the walls of the
excurrent canals; _Dermalia Autoderm[alia]_, microscleres in the dermal
membrane; _D. Hypoderm[alia]_, more deeply situated dermalia;
_Dictyonalia_, parenchymalia which become fused to form the skeletal
framework of Dictyonina; _Gastralia Autogastr[alia]_, microscleres in the
gastral membrane; _Gastralia Hypogastr[alia]_, more deeply situated
gastralia; _Parenchymalia Principalia_, main supporting spicules between
the chambers; _P. Comitalia_, slender diactine or triactine spicules
accompanying the last; _P. Intermedia_, microscleres between the P.
principalia; _Prostalia_, projecting spicules; _P. basalia_, rooting
spicules, from the base; _P. marginalia_, defensive spicules, round the
oscular rim; _P. pleuralia_, defensive spicules, from the sides. (From
Delage and Hérouard, after F. E. Schulze.)]
The deviations from this ground-plan of Hexactinellid structure are few and
simple. They are due to folding of the chamber-layer, or to variations in
the shape of the chambers, and to increasing fusion of the spicules to form
rigid skeletons. A simple condition of the chamber-layer, like that of the
young sponge of Fig. 89, {202}occurs also in some adult Hexactinellids,
_e.g._ in _Walteria_ of the Pacific Ocean (Fig. 90). Thus is represented in
this order the second type of canal system described among Calcarea. More
frequently, however, instead of forming a smooth sheet, the chamber-layer
grows out into a number of tubular diverticula, the cavities of which are
excurrent canals; these determine a corresponding number of incurrent
canals which lie between them. In this way there arises a canal system
resembling the third type of Calcarea. By still further pouching so as to
give secondary diverticula, opening into the first, a complicated canal
system is formed, as, for example, in _Euplectella suberea_.
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