The Cambridge natural history, Vol. 01 (of 10)Hickson, Sydney J. (Sydney John)
Science
The Cambridge natural history, Vol. 01 (of 10)
Hickson, Sydney J. (Sydney John)
Animals
Included in the Scyphozoa are the large jelly-fish found floating on the
sea or cast up on the beach on the British shores.
The Anthozoa include the Sea-anemones, nearly all the Stony Corals, the
Sea-fans, the Black Corals, the Dead-men's fingers (_Alcyonium_), the
Sea-pens, and the Precious Coral of commerce.
CLASS I. HYDROZOA
In this Class of Coelenterata two types of body-form may be found. In such
a genus as _Obelia_ there is a fixed branching colony of zooids, and each
zooid consists of a simple tubular body-wall composed of the two layers of
cells, the ectoderm and the {250}endoderm (Fig. 125), terminating distally
in a conical mound—the "hypostome"—which is perforated by the mouth and
surrounded by a crown of tentacles. This fixed colony, the "hydrosome,"
feeds and increases in size by gemmation, but does not produce sexual
cells. The hydrosome produces at a certain season of the year a number of
buds, which develop into small bell-like jelly-fish called the "Medusae,"
which swim away from the parent stock and produce the sexual cells. The
Medusa (Fig. 126) consists of a delicate dome-shaped contractile bell,
perforated by radial canals and fringed with tentacles; and from its centre
there depends, like the clapper of a bell, a tubular process, the
manubrium, which bears the mouth at its extremity. This free-swimming
sexual stage in the life-history of _Obelia_ is called the "medusome."
It is difficult to determine whether, in the evolution of the Hydrozoa, the
hydrosome preceded the medusome or _vice versâ_. By some authors the
medusome is regarded as a specially modified sexual individual of the
hydrosome colony. By others the medusome is regarded as the typical adult
Hydrozoon form, and the zooids of the hydrosome as nutritive individuals
arrested in their development to give support to it. Whatever may be the
right interpretation of the facts, however, it is found that in some forms
the medusome stage is more or less degenerate and the hydrosome is
predominant, whereas in others the hydrosome is degenerate or inconspicuous
and the medusome is predominant. Finally, in some cases there are no
traces, even in development, of a medusome stage, and the life-history is
completed in the hydrosome, while in others the hydrosome stages are lost
and the life-history is completed in the medusome.
If a conspicuous hydrosome stage is represented by H, a conspicuous
medusome stage by M, an inconspicuous or degenerate hydrosome stage by h,
an inconspicuous or degenerate medusome stage by m, and the fertilised ovum
by O, the life-histories of the Hydrozoa may be represented by the
following formulæ:—
1. O — H — O (_Hydra_)
2. O — H — m — O (_Sertularia_)
3. O — H — M — O (_Obelia_)
4. O — h — M — O (_Liriope_)
5. O — M — O (_Geryonia_)
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