The Cambridge natural history, Vol. 01 (of 10)Hickson, Sydney J. (Sydney John)
Science
The Cambridge natural history, Vol. 01 (of 10)
Hickson, Sydney J. (Sydney John)
Animals
In the Madreporaria a very elaborate skeleton of carbonate of lime is
formed. In the solitary forms it consists of a cup-shaped outer covering
for the base and column of the zooid called the "theca," of a series of
radial vertical walls or "septa" projecting into the intermesenteric
chambers carrying the endodermal lining of the coelenteric cavity with
them, and in some cases a pillar, the "columella," or a series of smaller
pillars, the "pali" projecting upwards from the centre of the base of the
{371}theca towards the stomodaeum. In the colonial forms the theca of the
individual zooids is continuous with a common colonial skeleton called the
"coenosteum." This is solid in the Imperforate corals, and it supports at
the surface only a thin lamina of canals and superficial ectoderm. In the
Perforate corals, however, the coenosteum envelopes and surrounds the
canals during its formation, and thereby remains perforated by a network of
fine channels. In the colonial Madreporaria the skeletal cups which support
and protect the zooids are called the "calices."
The skeleton of the Antipathidea is of a different nature. It is composed
of a horny substance allied to keratin. When it is old and thick, it
usually has a polished black appearance, and is commonly termed "black
coral." The surface of this kind of coral is ornamented with thorny or
spiny projections, but it is never perforated by calices or canal systems.
It forms a solid axis for the branches of the corals, and all the soft
parts of the zooids and coenosarc are superficial to it.
It was formerly considered that this type of coral, which shows no trace of
the shape and form of the living organisms that produce it, is of a
different character to the calcareous skeleton which exhibits calices,
septa, pores, and other evidence of the living organism, and it was called
a "sclerobase" to distinguish it from the "scleroderm" of the Madreporaria.
It is now known that both the sclerobasic skeleton and the sclerodermic
skeleton are products of the ectoderm, and consequently these expressions
are no longer in general use.
ASEXUAL REPRODUCTION in the Zoantharia may be effected by continuous or
discontinuous fission or gemmation.
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