The Cambridge natural history, Vol. 01 (of 10)Hickson, Sydney J. (Sydney John)
Science
The Cambridge natural history, Vol. 01 (of 10)
Hickson, Sydney J. (Sydney John)
Animals
Many secrete a stalk at the hinder end: by "continuous" formation of this,
without rupture at fission, a branching colony is formed (_Polyoeca_). This
stalk may have a varying consistency. In _Anthophysa_ (Fig. 37, 13) it
appears to be due to the welding of excrementitious particles voided at the
hinder end of the body with a gelatinous excretion; but the division of the
stalk is here occasional or intermittent, so that the cells are found in
tufts at the apex of the branches. A corresponding secretion, gelatinous or
chitinous, around the body of the cell forms a cup or "theca," within which
the cell lies quite free or sticking to it by its surface, or attached to
it by a rigid or contractile thread. The theca, again, may assume the form
of a mere gelatinous mass in which the cell-bodies may be completely
plunged, so that only the flagella protrude, as in Volvocidae,
_Proterospongia_ (Fig. 75, p. 182), and _Rhipidodendron_ (Fig. 37, 14).
Often this jelly assumes the form of a fan (_Phalansterium_), the branching
tubes of which it is composed lying for some way alongside, and ultimately
diverging. In _Hydrurus_, the branching jelly assumes the form of a
branching Confervoid.[117]
The cell-body may be bounded by an ill-defined plasmatic layer in
Chrysomonadaceae and some Protomastigaceae,[118] or it may form a plasmatic
membrane or "pellicle," sometimes very firm and tough, or striated as in
Euglenaceae, or it may have a separate "cuticle" (in the holophytic species
formed of cellulose), or even a bivalve or multivalve shell of distinct
plates, hinged or overlapping (_Cryptoglena_, _Phacotus_, Dinoflagellates).
The wall of the {114}Coccolithophoridae, a family of Chrysomonadaceae, is
strengthened by embedded calcareous spicules ("coccoliths," "cyatholiths,"
"rhabdoliths"), which in the most complex forms (cyatholiths) are like a
shirt-stud, traversed by a tube passing through the stem and opening at
both ends. These organisms[119] constitute a large proportion of the
plankton; the spicules isolated, or in their original state of aggregation
("coccospheres," "rhabdospheres"), enter largely into the composition of
deep-sea calcareous oozes. They occur fossil from Cambrian times (Potsdam
sandstone of Michigan and Canada), and are in some strata extremely
abundant, 800,000 occurring to the mm. cube in an Eocene marl.
The Silicoflagellates have siliceous skeletons resembling that of many
_Radiolaria_, to which they were referred until the living organism was
described (see pp. 79, 86 f.).
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