The Cambridge natural history, Vol. 02 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 02 (of 10)
Beddard, Frank E. (Frank Evers)
Animals
(i.) CHAETOSOMATIDAE.—This family includes three genera: _Chaetosoma_,
_Rhabdogaster_, and _Tristicochaeta_. According to Metschnikoff,[198]
although they are not true Nematodes, they have a great likeness to the
group. He distinguishes them from the swimming members of the group as
"creeping Nematoda." _Chaetosoma_, of which two species are known, _C.
ophicephalum_ and _C. claparedii_, has a head distinct from the body (Fig.
79). The mouth is at the anterior end, surrounded by a double semicircle of
movable spicules; the whole body is covered by fine hairs, and on the
ventral surface, just in front of the anus, is a double row of about
fifteen cylindrical projections, by whose agency the animal creeps. The
female _C. claparedii_ is 1.5 mm. long, the male 1.14 mm. They were found
creeping about on sea-weeds in the neighbourhood of Salerno.
[Illustration: FIG. 79.—Mature female of _Chaetosoma claparedii_ Metschni.,
× 57. (From Metschnikoff.) _a_, Oesophagus; _b_, intestine; _c_, anus; _d_,
ovary; _e_, generative pore; _f_, ventral bristles.]
The genus _Tristicochaeta_[199] differs from the foregoing in having three
rows of locomotor projections instead of two.
[Illustration: FIG. 80.—_Tristicochaeta inarimense_ Panceri, in one of its
most usual positions, showing the triple row of ventral bristles, × 100.
(From Panceri.)]
_Rhabdogaster_ has no head distinct from the body, though the anterior part
of the body is swollen. A second swelling occurs, as is also the case with
_Chaetosoma_, in the region of the opening of the genital ducts. The female
in _Rh. cygnoides_ attains a length of 0.36 mm. In this genus the hairs are
confined to {159}the dorsal middle line. The locomotor projections are
hooked, and are much finer than those of _Chaetosoma_, and they are
situated farther forward than in the last-named genus. _Rhabdogaster_
occurs in the same surroundings as _Chaetosoma_. _Ch. ophicephalum_ is
recorded from the English Channel.
(ii.) DESMOSCOLECIDAE.—The members of this family are minute, and are
characterised by the presence of well-marked ridges which surround the body
and give it an appearance of segmentation. The head, which is somewhat
swollen, bears four bristles, and single pairs are borne by a certain
number of the ridges, some on the dorsal and some on the ventral surface.
These hairs can be moved independently of one another. Two red eye-spots
are described between the fourth and fifth rings. The sexes are distinct,
and the internal organs generally have a marked resemblance to those of the
true Nematoda. The _Desmoscolecidae_ move by looping their bodies after the
manner of the Geometrid caterpillars, as well as by creeping with their
bristles. The genus contains numerous species[200]: _D. minutus_ Clap.
(English Channel), _D. nematoides_ Greef, _D. adelphus_ Greef, _D.
chaetogaster_ Greef, _D. elongatus_ Panceri, and _D. lanuginosa_ Panceri.
They are exclusively marine.
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