The Cambridge natural history, Vol. 02 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 02 (of 10)
Beddard, Frank E. (Frank Evers)
Animals
They exist in incredible numbers; Grassi describes the surface of the sea
at Messina on certain days as being literally covered with them, and they
must form the food supply of numerous animals which prey upon the pelagic
fauna. The immense number of individuals is probably accounted for to some
extent by the fact that they lay eggs all the year round, and pass
{191}through a very short and rapid development. They are not known to be
phosphorescent.
CLASSIFICATION.—The features of the Chaetognatha which have most systematic
value are the size of the adult, the relations of the length to the
breadth, and of the three divisions to one another; the size, number, and
position of the lateral fins, and of the hooks and spines on the head; the
thickness of the epidermis, and the structure of the olfactory organ; and,
finally, the form of the reproductive organs.
Strodtmann,[229] who gives the latest and most complete account of the
species of Chaetognatha, arranges them under three genera, which he
characterises as follows:—
(i.) _Sagitta_ Slabber.—Two pairs of lateral fins, two rows of spines on
the head. The lateral thickening of the epidermis absent or insignificant.
Under this genus are included nine definite species and five others—_S.
gracilis_ Verrill, _S. elegans_ Verrill, _S. darwini_ Grassi, _S. diptera_
d'Orbigny, and _S. triptera_ d'Orbigny—whose position, owing to the
inadequacy of their description, is of doubtful validity.
[Illustration: FIG. 104.—_Sagitta hexaptera._ Ventral view. × 4. (From
Hertwig.) _a_, Mouth; _b_, hooks; _c_, anterior septum; _d_, alimentary
canal; _e_, commissure from the brain to the ventral ganglion; _f_, ventral
ganglion; _g_, ovary; _h_, oviduct; _i_, posterior septum; _j_, testis;
_k_, vesicula seminalis.]
The distribution of the other species may be mentioned. _S. hexaptera_ is
the largest Chaetognath known, and reaches in the adult stage a length of 7
cm. It is very widely distributed, being found in practically all the
temperate and warm seas, usually at the surface of the water, though at
times it is found at a depth of one metre, or even deeper. _S. lyra_,
Mediterranean, very rare. _S. tricuspidata_, widely distributed. _S.
magna_, Mediterranean and Madeiran, living at the surface. _S. bipunctata_,
the most frequently described form, smaller than the preceding species, 1-2
{192}cm. in length, widely distributed, and as a rule living near the coast
line. _S. serratodentata_, Mediterranean. _S. enflata_, on the surface of
the sea, Mediterranean and Madeiran. _S. minima_, a very small species, 1
cm. in length, Mediterranean. _S. falcidens_, Atlantic, off the coast of
New Jersey.
(ii.) _Krohnia_ Langerhans.—A single lateral fin extending on to both trunk
and tail segment, no lateral epidermal extensions behind the head, only one
row of spines on the head. Trunk longer than the tail.
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