The Cambridge natural history, Vol. 02 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 02 (of 10)
Beddard, Frank E. (Frank Evers)
Animals
[Illustration: FIG. 126.—Diagrams to illustrate the action of the
Chaetopodan "introvert." (From Lang.) A shows the apparatus at rest; the
mouth leads into the buccal cavity (_B_), with paragnaths in its wall; _P_,
the pharynx, with jaws at its anterior end; _c_, brain; _p_, protractor
muscles; _r_, retractors. B, the pharynx has been brought forward or
protruded by the eversion, or turning inside out, of the buccal region, so
that the jaws (_j_) now lie some way in front of the head, which is
represented by the brain.]
The buccal region is lined with chitin, which is specially thickened at
certain definite spots, forming small "denticles" or "paragnaths" (Fig.
125), which have a different arrangement in the various species.
The cavity of the pharynx is narrow and the walls thick and muscular; each
side wall carries a large, dark, chitinous "jaw" (Fig. 127, J), which is
hollow at the base, into which the muscles serving to move it are inserted,
whilst the apex is solid, curved, and more or less notched. These two great
jaws are used not only for tearing prey, but for seizing it; for when the
pharynx is entirely protruded the two jaws are wide apart, and when
retraction takes place they come together and grasp the prey.
Eversion of the apparatus is partly effected by protractor muscles (Fig.
126, A, _p_) and partly by the pressure of the coelomic fluid, compressed
by the muscles of the body-wall; the eversion is stopped at a certain stage
by a sheet of muscular tissue or "diaphragm" (Fig. 127, _diaph_) inserted
round the buccal region and attached to the body-wall in the second
segment. The introversion is effected partly by the contraction of this
diaphragm and partly by the action of powerful retractor muscles (Fig. 126,
_r_) inserted into the hinder end of the pharynx and passing to the
body-wall (these are removed in Fig. 127). The {251}movement of the jaws
themselves and of the wall of the apparatus is due to other muscles.
The _oesophagus_ is quite short; into it opens a pair of sacculated
diverticula or glands. Then follows the _intestine_, which extends through
the rest of the body as a thin-walled tube, slightly dilated at the
insertion of the septa.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account