The Cambridge natural history, Vol. 02 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 02 (of 10)
Beddard, Frank E. (Frank Evers)
Animals
All the many hundreds of species of Polychaetes are marine, with a very few
exceptions, which have been in recent years recorded from fresh (_i.e._
drinkable) water, viz. a species of _Nereis_ from a lake in Mingrelia,
another _Nereis_ and a _Lumbriconereis_ from running water in
Trinidad,[340] a Sabellid, _Manayunkia speciosa_,[341] from Philadelphia;
and another Sabellid, _Coabangia_,[342] from fresh water at Tonquin, which
lives in borings in shells of _Melania_; and it is by no means improbable
that other fresh-water Polychaetes exist in Lake Tanganyika in Africa,
where a Medusa has recently been discovered.
In brackish water of various densities many Polychaetes live; _Arenicola_
especially is regardless of the character of the medium, and _Nereis
diversicolor_ appears to withstand considerable admixture of fresh water.
The majority of the Polychaetes occur "inshore," that is, between
tide-marks and in shallow water down to 20 fathoms; but they occur at all
depths more or less abundantly, and some have been dredged from depths of
more than 3000 fathoms.
The nature of the soil composing the shore has a good deal to do with the
number of worms to be found there; thus in calcareous districts they are
fewer than in places where harder rocks, {285}such as granite, form the
shore line, for the chalk or limestone wears away more quickly, and exposes
to destruction the worms which may have sheltered in its crevices: further,
it does not give so permanent a place of attachment to seaweeds, on which
many Polychaetes feed. The calcareous rocks, too, are more likely to be
traversed by springs of fresh water, which is not to the taste of the
worms. The sand resulting from the destruction of the rocks, whether hard
or soft, is of itself unsuitable to the majority of worms, which are most
abundant where mud containing decaying vegetable matter is mixed with the
sand: this, which gives a firmer consistency to the soil, so that the
burrows retain their form better, supplies food for the burrowers.
GENERAL HABITS.—The division of the Polychaetes into the "Errantia" or
free-swimming and wandering forms, and "Sedentaria" or tubicolous and
sedentary forms, is a misleading mode of classification, for as a matter of
fact only a comparatively few forms are really free-swimming throughout
life; the majority, even if they do not form definite tubes, burrow
galleries for themselves in the soil, and these burrows are in many cases
only rarely left; this is true of both groups. Amongst the "errant"
Polychaetes nearly all the Eunicidae secrete a parchment-like tube, and
some Polynoids form mud tubes. Among the "sedentary worms" there are forms
which merely burrow; while _Myxicola_ readily leaves its gelatinous tube
and swims freely; _Pectinaria_ carries its house with it as it moves about,
and _Polycirrus_, a Terebellid, does not form any tube at all.
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