The Cambridge natural history, Vol. 02 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 02 (of 10)
Beddard, Frank E. (Frank Evers)
Animals
the remaining segments; and along the outer edge of each are some ten
bundles of chaetae, and along the hinder edge some five or six bundles,
representing as many segments. Thus the worm consists of about thirty
segments whose outlines are nearly obliterated (as in the _Chlorhaemidae_),
and whose chaetae, except those which are specially developed, are
disappearing: while posteriorly a great shortening of the body brings the
bundles close together. A continuation of this process, involving a further
disappearance of chaetae, leads readily to the condition met with in
_Echiurus_, one of the chaetigerous Gephyrea. Internally, further evidence
of the relation between _Sternaspis_ and the Chlorhaemids with the Gephyrea
is afforded by the absence of septa, by the coiled alimentary canal, and by
the presence of a single pair of nephridia, which in the latter group act
both as excretory organs and as genital ducts.
BRANCH B. CRYPTOCEPHALA.
SUB-ORDER 1. SABELLIFORMIA.[392]
FAM. 1. _Sabellidae._—The branchial crown consists of a usually
considerable number of filaments arising from a semicircular base. The
peristomium may be reflexed to form a collar, {337}which is frequently
notched, so that a lateral and a ventral lobe on each side may be
distinguished (Fig. 133, B, p. 261). The thorax consists of nine segments,
and is provided with ventral "gland shields," which are continued along the
abdomen, where they are subdivided into two by a ciliated "faecal groove,"
which sometimes bends to one side on reaching the thorax, and may extend
forwards along the dorsal surface to the head: this groove serves to carry
the faeces out of the tube. The gill filaments are always provided with
secondary processes, and may be provided with compound eyes.[393] The tubes
of the Sabellidae are always of fine mud or of sand.
_Sabella pavonia_ Sav. is about 10 to 12 inches long and about ¼ inch
across; the tube of fine mud is considerably longer and embedded deeply in
the mud, with its free end projecting to some 2 or more inches, where it
serves for the attachment of seaweed, Polyzoa, Hydrozoa, etc. The colour of
the animal is orange-brown; the gills, which are about 1½ inches long, are
green (due to contained blood) marked with more or less extensive brown or
purple-brown spots, which may even hide the green tint. There is a pair of
dark filaments arising between the dorsalmost gill filaments, which have
been erroneously regarded as "prostomial tentacles"; they are, in fact,
prolongations of a peculiar membrane or lip round the base of the gills,
which bounds a groove leading to the mouth. These lip-processes (Fig. 133,
B, _l_) occur in other Sabellids. Atlantic, North Sea, and Mediterranean.
[Illustration: FIG. 182.—A, A gill filament of _Branchiomma_; B, of
_Dasychone_. _a_, Axis; _f_, secondary filaments; _o_, eye; _x_, dorsal
appendices.]
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