The Cambridge natural history, Vol. 02 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 02 (of 10)
Beddard, Frank E. (Frank Evers)
Animals
NERVOUS SYSTEM.[131]—The brain is composed of two ganglionic masses (Fig.
53, _n.g_) lying at the anterior end of the body, one on each side of the
proboscis, and united by commissures passing round it (Fig. 55, _d.c_ and
_v.c_). Each ganglionic mass is often partially divided into a dorsal and
ventral lobe (_n.g.d_ and _n.g.v_). From the brain a pair of cords pass off
backwards along the sides of the body (_n.c_); these cords, which have no
ganglionic swellings, in some forms unite with one another above the anus.
Anteriorly nerves are given off from the brain to the eyes and front part
of the head (_a.n_). A nerve to the proboscis is given off from the
commissure which unites the two halves of the brain dorsal to the proboscis
(_d.n_).
[Illustration: FIG. 54.—Diagrammatic representation of the proboscis, (A)
in the retracted condition, (B) in the everted condition. _g.p_, Glandular
portion of the proboscis; _m_, muscle attaching the proboscis to its
sheath; _m.p_, muscular portion of the proboscis; _p.p_ in A, proboscis
pore; _p.p_ in B represents the position of the proboscis pore in the
retracted condition of the proboscis; _p.s_, proboscis sheath.]
In two out of the three groups into which the Nemertines are divided, the
lateral nerve-cords are in connexion with a network or plexus of nerves
lying between the muscular layers of the body-wall (Fig. 52, _n.l_), and in
some forms constituting a comparatively thick layer. In these two groups
there are no definite {106}nerve branches except the anterior ones to the
head. In the third group of Nemertines the lateral nerve-cords lie within
the muscular layers of the body-wall, and in this case paired nerve
branches are given off at definite intervals throughout the whole length of
the body. These branches divide up among the organs to which they pass, and
no nerve plexus is present.
The lateral cords vary in position in different cases. Sometimes they lie
laterally, at others the cords tend to approximate to one another in the
median dorsal or in the median ventral line, though in every case they
remain distinctly separated.
SENSE ORGANS.—Sense organs are usually present in the form of eyes arranged
at the sides of the head (Fig. 51, _e_), sometimes as a single pair and
sometimes in one or more groups on each side. The structure of the eyes
varies from a simple pigment spot to an organ which receives a special
nerve-supply from the brain, and possesses a refracting body answering to a
lens, and behind this a pigment layer and a layer of rods. Some forms are
devoid of all traces of eyes.
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