The Cambridge natural history, Vol. 03 (of 10)Cooke, A. H. (Alfred Hands)
Science
The Cambridge natural history, Vol. 03 (of 10)
Cooke, A. H. (Alfred Hands)
Animals
Following first the oviduct, we find that it soon widens into the
_vagina_ (V.), which is furnished with a pair of _mucous glands_
(M.G.), one on each side. These are much branched, and resemble little
bunches of whitish seaweed. A little above the mucous glands a long
tube diverges from the vagina, which is furnished with a produced
_coecum_ (C.) and a pouch, the _spermatheca_ (SP.) at the extreme
end. In this pouch, and in the duct leading to it, is stored the
spermatophore received in union with another snail. Just below the
mucous glands the vagina is joined by the _dart sac_ (D.S.), which is
more fully described below. Finally, at its lower end the vagina unites
with the penis sac at a point just posterior to the common orifice.
Returning now to the male organs, we find that the _vas deferens_
is the continuation of the male portion of the hermaphrodite duct,
after its final separation from the female portion. It passes under
the retractor muscle of the upper right tentacle, which has been cut
away in the specimen figured, to dissect it out. Just before the vas
deferens widens into the penis sac, it branches off into a long and
tapering tube, the _flagellum_, in which the spermatozoa are stored and
become massed together in the long packet known as the _spermatophore_.
The _penis sac_ (P.S.) is the continuation of the vas deferens beyond
the point at which the flagellum diverges. It joins the vagina at
its extreme anterior end, uniting with it to form the common genital
aperture, which cannot be exactly represented in the figure. The
_penis_ itself lies in the interior of the penis sac, and is a rather
long muscular tube which is protruded during union, but at other times
remains retracted within the sac.
In the Helicidae generally, the form of the generative organs varies
with each separate species, sometimes merely as regards the size of
the different parts, at others in the direction of greater simplicity
or complication. The mucous glands may be absent, and the flagellum
greatly reduced in size, or absent altogether.
=The Dart Sac.=--A remarkable part of the reproductive system in many
of the true Helicidae is the so-called _dart_, _Liebespfeil_, or _telum
veneris_. It consists of ‘a straight, or curved, sometimes slightly
twisted tubular shaft of carbonate of lime, tapering to a fine point
above, and enlarging gradually, more often somewhat abruptly, to
the base.’ The sides of the shaft are sometimes furnished with two
or more blades; these are apparently not for cutting purposes, but
simply to brace the stem. The dart is contained in a dart sac, which
is attached as a sort of pocket to the vagina, at no great distance
from its orifice. There are four different forms of sac. It may be
single or double, and each of these divisions may be bilobed, each lobe
containing one dart at a time. In _Helix aspersa_ the dart is about
5/16 in. in length, and ⅛ in. in breadth at its base (see Fig. 54).
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