The Cambridge natural history, Vol. 03 (of 10)Cooke, A. H. (Alfred Hands)
Science
The Cambridge natural history, Vol. 03 (of 10)
Cooke, A. H. (Alfred Hands)
Animals
=Ocelli of Pecten.=--In _Pecten_ and _Spondylus_ the _ocelli_ are
remarkably large and prominent, shining like precious stones, and are
placed along the two edges of the mantle so as to receive the light
when the shell gapes (Fig. 93). In _Pecten opercularis_, _jacobaeus_,
and _maximus_ their number varies from 80 to 120. In _Spondylus
gaederopus_, a very inequivalve shell, 60 have been counted on the
right or fixed valve, and 90 on the left or upper valve. Each ocellus
is connected, by means of its optic nerve, with the large circumpalleal
nerve, and so with the branchial ganglion. They possess a cornea,
lens, choroidea, and optic nerve, and, according to Hickson,[298] bear
a considerable resemblance to the vertebrate type of eye. In spite
of this, the power of vision in these genera does not appear at all
superior to that of other Pelecypoda.
[Illustration: FIG. 93.--_Pecten opercularis_ L., showing the
_ocelli_ on the two edges of the mantle.]
[Illustration: FIG. 94.--Compound eyes (_c.e_) of _Arca barbata_
L.; _m.l_, mantle fold; _omm_, ommatidia. (After Patten.)]
According to the elaborate investigations of Patten, the ‘eyes’ in
_Arca_ occur upon the middle or ‘ophthalmic’ fold of the mantle-edge,
which is thickened at the end to admit of their reception. Along this
is ranged a row of dark brown spots of various sizes, which are larger
at the anterior and posterior ends of the mantle-edge, but smaller and
more numerous towards the middle. These brown spots, or ‘eyes,’ are
many of them compound, being made up of the fusion of a number of
ommatidia (from 10 to 80) into one large round eye, which is generally
elevated above the surface of the surrounding epithelium. Sometimes
these eyes themselves tend to fuse together. In one specimen of _Arca
Noae_, 133 of these faceted eyes were counted in one mantle border, and
102 in the other.
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