The Cambridge natural history, Vol. 03 (of 10)Cooke, A. H. (Alfred Hands)
Science
The Cambridge natural history, Vol. 03 (of 10)
Cooke, A. H. (Alfred Hands)
Animals
=Pelecypoda.=--The nervous system in the Pelecypoda is the simplest
type in which well-marked ganglionic centres occur. The ganglia are
few, symmetrically placed, and are usually at a considerable distance
apart. There are, as a rule, three distinct pairs of ganglia, the
cerebral (cerebro-pleural), pedal, and visceral. The cerebral are
formed by the fusion of the cerebral and pleural ganglia, which,
however, in some cases (Protobranchiata) continue distinct.[317] They
lie above or on each side of the mouth, united by a commissure of
varying length. Another pair of commissures unites them with the pedal
ganglia, which are placed at the base of the foot, and are usually
very close together, sometimes (as in _Anodonta_) becoming partially
fused. The length of these commissures depends upon the distance
between mouth and foot; thus they are very long in _Mya_ and _Modiola_,
and very short in _Pecten_. In cases where the foot is rudimentary or
becomes aborted through disuse (_e.g._ _Ostrea_), the pedal ganglia
may dwindle or disappear altogether. The visceral ganglia are on the
ventral side of the posterior adductor muscle, beneath the rectum,
and innervate the branchiae, osphradia, and the whole of the visceral
sac. A pair of cerebro-visceral commissures traverses the base of the
foot, surrounding it with a comparatively short loop (compare Fig. 106,
_c.v.c´_), while a long commissure, which runs round the entire edge
of the mantle, and supplies branching nerves to the mantle border and
siphons (Fig. 106, _c.v.c_), may also connect the visceral and cerebral
ganglia.
[Illustration: FIG. 105.--Nervous system of Pelecypoda: _A_,
_Teredo_; _B_, _Anodonta_; _C_, _Pecten_; _a_, _a_, cerebral
ganglia; _b_, pedal ganglia; _c_, visceral ganglia. (After
Gegenbaur.)]
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