The Cambridge natural history, Vol. 03 (of 10)Cooke, A. H. (Alfred Hands)
Science
The Cambridge natural history, Vol. 03 (of 10)
Cooke, A. H. (Alfred Hands)
Animals
It must not be forgotten, however, that a simple repetition of the same
tooth, whether lateral or marginal, does not necessarily constitute
an important characteristic, nor does the presence or absence of a
central tooth. In most of the cases mentioned above, the difference in
the number of laterals and marginals is due to the multiplication of
identical forms, while the central tooth, when present, is often a mere
plate or narrow block without cusps, whose presence or absence makes
little difference to the character of the radula as a whole.
There appear to be three well-marked types of radula among the
Opisthobranchiata.
(_a_) Radula with a single strong central tooth, rows few. This form
is characteristic of the Aeolididae, Fionidae, Glaucidae, Dotoidae,
Hermaeidae, Elysiidae (Fig. 135), and Limapontiidae. In the Aeolididae
it is sometimes accompanied by a single lateral. The same type occurs
in _Oxynoe_, and in _Lobiger_ (= _Lophocercus_).
(_b_) Radula with the first lateral very strongly developed. This type
may take the form of (1) a single lateral, no central or marginals,
_e.g._ _Onchidoris_, _Scaphander_ (Fig. 137, A), _Philine_ (certain
species), _Ringicula_, or (2) first lateral strongly developed, and
repeated in succeeding laterals (2–6) on a smaller scale, _e.g._
_Philine_ (certain species). A few marginals are sometimes added,
_e.g._ in _Polycera_, _Lamellidoris_ (where there is a degraded central
tooth, Fig. 137, B), _Idalia_, and _Ancula_.
[Illustration: FIG. 135.--Radula of _Elysia viridis_ Mont. × 40.
Type (_a_).]
[Illustration: FIG. 136.--Portion of the radula of _Gadinia
peruviana_ Sowb., Chili. × 250. Type (_c_).]
(_c_) Radula with an indefinite number of marginals, laterals (if
present) merging into marginals, central tooth present or absent,
inconspicuous, teeth all very small. This type of radula, among the
Nudibranchiata, is characteristic of certain sub-genera of _Doris_
(_e.g._ _Chromodoris_, _Aphelodoris_, _Casella_, _Centrodoris_), of
_Hypobranchiaea_ and _Pleurophyllidia_; among the Tectibranchiata,
of _Actaeon_, many of the Bullidae, _Aplustrum_, the Aplysiidae,
_Pleurobranchus_, _Umbrella_ and _Gadinia_ (Figs. 136 and 137, C).
In the _Pteropoda_ there are two types of radula. The Gymnosomata,
which are in the main carnivorous, possess a radula with a varying
number (4–12) of sickle-shaped marginals, central tooth present or
absent. In the Thecosomata, which feed on a vegetable diet, there are
never more than three teeth, a central and a marginal on each side;
teeth more or less cusped on a square base.
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