The Cambridge natural history, Vol. 03 (of 10)Cooke, A. H. (Alfred Hands)
Science
The Cambridge natural history, Vol. 03 (of 10)
Cooke, A. H. (Alfred Hands)
Animals
(_d_) _Province of the Great Lakes._--The Mollusca of the four great
lakes of Eastern Central Africa--Lakes Albert Nyanza (Luta Nzige,
2720 ft.), Victoria Nyanza (Oukéréwé, 3700 ft.), Nyassa (1520 ft.),
and Tanganyika (2800 ft.)--are well known, and supply an interesting
problem in distribution. Those of the three first mentioned lakes
differ in no way from the rest of tropical Africa, but the Mollusca
of Tanganyika include, in addition to the ordinary African element,
a number of peculiar operculate genera, belonging principally to the
Melaniidae and Hydrobiidae. Several of these possess a solidity of form
and compactness of structure which is unusual in fresh-water genera,
and has led to the belief, among some authorities, that they are the
direct descendants of marine species, and that Tanganyika represents
an ancient marine area. This view appears untenable. The Victoria
Nyanza and Nyassa are part of the same system as Tanganyika, and it
is not easy to see how, if Tanganyika were once an arm of the sea,
they were not equally so, especially as they are several hundred miles
nearer the Indian Ocean as at present defined. Nor, as will be seen
from the figures given above, is there anything in the altitudes which
would make us expect anything exceptional in Tanganyika. The similar
case of L. Baikal must be compared (p. 290), where again a number of
specialised forms of _Hydrobia_ occur.
Of the genera concerned, _Paramelania_ and _Nassopsis_ are forms of
Melaniidae; _Tiphobia_ (Fig. 219), which is allied to _Paludomus_, is a
compact shell with angulated spinose whorls; _Lacunopsis_, _Ponsonbya_,
_Limnotrochus_, and _Tanganyicia_ are probably forms of _Lithoglyphus_,
some, as their names denote, being of decidedly marine facies;
_Syrnolopsis_ and _Turbonilla_ (?) look like Pyramidellidae, _Horea_
and _Reymondia_ like _Rissoina_; _Bourguignatia_ appears to belong
to _Vivipara_, with which has now been merged the genus _Neothauma_.
Recently discovered forms from the adjacent L. Mweru are evidently of
kindred origin.
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