The Cambridge natural history, Vol. 03 (of 10)Cooke, A. H. (Alfred Hands)
Science
The Cambridge natural history, Vol. 03 (of 10)
Cooke, A. H. (Alfred Hands)
Animals
On _Tunicata_.--_Lamellaria_ deposits its eggs and lives on an Ascidian
(_Leptoclinum_), and the common _Modiolaria marmorata_ lives in
colonies imbedded in the test of _Ascidia mentula_ and other simple
Ascidians.
[Illustration: FIG. 31.--Two species of _Eulima_: =A= is sessile
on the skin of a Holothurian, through which it plunges its
sucking proboscis (_Pr_); =B= creeps freely in the stomach of a
Holothurian. (After K. Semper.)]
Special points of interest with regard to parasitic Mollusca relate
to (1) _Colour_. This is in most cases absent, the shell being of
a uniform hyaline or milky white. This may be due, in the case of
the endo-parasitic forms, to absence of light, and possibly, in
those living outside their host, to some deficiency in the nutritive
material. A colourless shell is not necessarily protective, for
though a transparent shell might evade detection, a milk-white hue
would probably be conspicuous. (2) _Modifications of structure._
These are in many cases considerable. _Entoconcha_ and _Entocolax_
have no respiratory or circulatory organs, and no known nervous
system; _Thyca_ and certain _Stilifer_ possess a curious suctorial
apparatus; the foot in many cases has aborted, since the necessity for
locomotion is reduced to a minimum, and its place is supplied by an
enormous development of the proboscis, which enables the creature to
provide itself with nutriment without shifting its position. K. Semper
notices a case where a _Eulima_, whose habitat is the stomach of a
Holothurian, retains the foot unmodified, while a species occurring
on the outer skin, but provided with a long proboscis, has lost its
foot altogether.[177] Special provision for holding on is noticed in
certain cases, reminding us of similar provision in human parasites.
Eyes are frequently, but not always wanting, even in endo-parasitic
forms. A specially interesting modification of structure occurs in
(3) the _Radula_ or ribbon-shaped arrangement of the teeth. In most
cases of parasitism (_Eulima_, _Stilifer_, _Odostomia_, _Entoconcha_,
_Entocolax_, _Magilus_, _Coralliophila_, _Leptoconcha_) it is absent
altogether. In _Ovula_ and _Pedicularia_, genera which are in all
other respects closely allied to _Cypraea_, the radula exhibits marked
differences from the typical radula of the Cypraeidae. The formula
(3·1·3) remains the same, but the laterals are greatly produced
and become fimbriated, sometimes at the extremity only, sometimes
along the whole length. A very similar modification occurs in the
radula of _Sistrum spectrum_ Reeve, a species which is known to live
parasitically on one of the branching corals. Here the laterals differ
from those of the typical _Purpuridae_ in being very long and curved at
the extremity. The general effect of these modifications appears to be
the production of a radula rather of the type of the vegetable-feeding
_Trochidae_, which may perhaps be regarded as a link in the chain of
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