The Cambridge natural history, Vol. 03 (of 10)Cooke, A. H. (Alfred Hands)
Science
The Cambridge natural history, Vol. 03 (of 10)
Cooke, A. H. (Alfred Hands)
Animals
The immediate neighbourhood of the sea appears frequently to have the
effect of dwarfing land Mollusca. Thus the var. _conoidea_ of _Helix
aspersa_, which is small, conical, with a compressed mouth, occurs
‘on sandhills and cliffs at the seaside.’ The varieties _conica_ and
_nana_ of _Helix hispida_ are found ‘near the sea.’ _Helix virgata_
is exceedingly small in similar localities, and tends to become
unicoloured. _H. caperata_ var. _Gigaxii_, a small depressed form,
occurs at ‘Sandwich and Falmouth.’[193] Sometimes, however, the exact
opposite is the case, for _H. nemoralis_ var. _major_, which is
‘much larger’ than the type, occurs on ‘sandhills and downs’ and is
‘remarkably large in the I. of Arran, Co. Galway.’ The dwarf form of
_Limnaea peregra_ known as _maritima_ appears to be confined to the
neighbourhood of the sea.
Dwarfing of the shell seems frequently to be the result of an elevated
locality, not perhaps so much as the direct consequence of purer air
and less barometric pressure, as of changes in the character of the
food supply and in the humidity of the air. Several species of _Helix_
have a variety _minor_ which is characteristic of an Alpine habitat.
_Helix arbustorum_ var. _alpestris_, which is scarcely two-thirds the
size of the type, occurs on the Swiss Alps in the region of perpetual
snow. Sometimes a very slight elevation is sufficient to produce the
dwarfed form. At Tenby the type form of _Helix pisana_ is scattered in
countless numbers over the sandhills just above high-water mark. At the
extreme western end of these sandhills rises abruptly to a height of
over 100 feet the promontory known as Giltar Head, the vegetation of
which is entirely distinct from that of the burrows below. There is a
colony of _H. pisana_ at the end of Giltar, all of which are devoid of
the characteristic markings of the typical form, and most are dwarfed
and stunted in growth.
Occasionally the same variety will be found to be produced by
surroundings of very different nature. Thus the var. _alpestris_, of
_H. arbustorum_ mentioned above, besides being characteristic of high
Alpine localities, also occurs abundantly in low marshes at Hoddesdon
on the River Lea. _Helix pulchella_ var. _costata_, according to
Jeffreys, is found in dry and sandy places, often under loose stones
and bricks on walls, while other authorities have noticed it in wet and
dry localities quite indifferently.
Sometimes the production of a variety may be traced to the intrusion
of some other organism. According to Brot, nine-tenths of the _Limnaea
peregra_ inhabiting a certain pond near Geneva, were, during one
season, afflicted with a malformation of the base of the columella.
This deformity coincided with the appearance, in the same waters, of
extraordinary numbers of _Hydra viridis_. The next season, when the
_Hydra_ disappeared, the next generation of _Limnaea_ was found to
have resumed its normal form.
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