The Cambridge natural history, Vol. 04 (of 10)Warburton, Cecil
Science
The Cambridge natural history, Vol. 04 (of 10)
Warburton, Cecil
Animals
Although Trilobites are often found in abundance and in an excellent
state of preservation, it is only in very rare cases that anything is
seen of the ventral surface except the hypostome and the reflexed
borders of the cephalic shield, of the thoracic segments, and of the
pygidium. The usual absence of =appendages= is probably due to their
tenuity. Billings, in 1870, first obtained clear evidence of the
presence of pairs of appendages, in _Asaphus platycephalus_. Soon
afterwards Walcott[191] showed their existence in American specimens of
_Asaphus megistos_, _Calymene senaria_, and _Cheirurus pleurexacanthus_.
In the two latter species the appendages were found by cutting sections
of curled-up specimens obtained from the Trenton Limestone; 2200
examples were sliced, of which 270 showed evidence of the existence of
appendages. They were seen to be present on the head, thorax, and
pygidium; a ventral uncalcified cuticle with transverse arches was also
found. By means of sections of curled-up specimens it was difficult to
determine satisfactorily the form and position of the appendages.
Subsequently extended specimens of _Triarthrus_ (Fig. 142) and
_Trinucleus_, showing the ventral surface and appendages clearly, were
discovered in the Utica Slate (Ordovician) near Rome, New York. A full
account of the appendages in those specimens has been given by
Beecher.[192]
[Illustration:
FIG. 142.—_Triarthrus becki_, Green, × 2½. Utica Slate (Ordovician),
near Rome, New York. =A=, Ventral surface with appendages; _Ep_,
metastome; _Hy_, hypostome. =B=, second thoracic appendage; _en_,
endopodite; _ex_, exopodite, × 12. (After Beecher.)
]
In _Triarthrus_ each segment, except the anal, bears a pair of
appendages, all of which, except the first, are biramous. There are five
pairs of cephalic appendages; the first pair are attached at each side
of the hypostome, and have the structure of antennae, each consisting of
a single flagellum formed of short conical joints. The other cephalic
appendages increase in size successively. At present the second and
third pairs are not satisfactorily known, but appear to have been
similar to the fourth and fifth pairs. The second pair is attached at
the level of the posterior end of the hypostome. The fourth and fifth
pairs have large, triangular coxopodites which served as gnathobases,
their inner edges being denticulate; the endopodites consist of stout
joints; the exopodites are slender, and bear setae which are often
arranged in a fan-like manner.
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