The Cambridge natural history, Vol. 04 (of 10)Warburton, Cecil
Science
The Cambridge natural history, Vol. 04 (of 10)
Warburton, Cecil
Animals
A modification of Salter’s classification has been brought forward by
Beecher[200] who divides the Trilobita into three main groups:—
1. _Hypoparia._ Facial sutures at or near the margin, or ventral.
Compound eyes absent. This is equivalent to Salter’s Agnostini and
Ampycini with the addition of the Harpedidae.
2. _Opisthoparia._ Facial sutures extending from the posterior margin to
the front margin, but occasionally uniting in front of the glabella.
Eyes holochroal or prismatic, but sometimes absent. This comprises the
same families as Salter’s Asaphini with the exclusion of the Harpedidae
and Calymenidae.
3. _Proparia._ Facial sutures extending from the lateral margins, and
either cutting the anterior margin or uniting in front of the glabella.
Eyes holochroal or schizochroal; occasionally absent. This is equivalent
to Salter’s Phacopini with the addition of the Calymenidae.
In each of the groups proposed Beecher regards as the more primitive
forms those which possess characters similar to those of the early
larval stages, such as narrow free cheeks, the absence of compound eyes,
and a glabella which is broad in front and reaches the anterior margin
of the head.
The modifications introduced by Beecher can scarcely be regarded as
making Salter’s classification more natural. For instance, the
Agnostidae differ so much from all other families that, at present,
there is no evidence to show that they have any close phylogenetic
relationship with the Trinucleidae and Harpedidae. Further, the
Calymenidae, which Salter recognised as related to the Olenidae, have
been shown by the careful work of Professor Pompeckj[201] to have
descended from the latter family, and to have no genetic connexion with
the Phacopidae with which they are grouped by Beecher. Then in the
Trinucleidae the earliest genus, _Orometopus_[202] (Fig. 140, A),
possesses compound eyes and facial sutures which begin at the posterior
margin and unite in front of the glabella; so that, according to
Beecher’s classification, that genus would belong to the Opisthoparia,
whereas the later genera (_Trinucleus_, etc.) of the same family would
be placed in the Hypoparia. At present, therefore, the only
classification of Trilobites which can be adopted is a division into
families, of which a short account is given below.
[Illustration:
FIG. 146.—_Agnostus integer_, Beyr., × 8. Cambrian. (After Barrande.)
]
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account