The Cambridge natural history, Vol. 04 (of 10)Warburton, Cecil
Science
The Cambridge natural history, Vol. 04 (of 10)
Warburton, Cecil
Animals
convenient that each joint should have a separate name, but it must be
borne in mind that the tibia of different authors is not always the same
joint. Special attention must be directed to the three terminal joints,
which furnish highly important characteristics. The tibia (in Pocock’s
sense) is sometimes provided with a “tibial spur” at its lower distal
extremity. From the soft arthrodial membrane between the protarsus and
tarsus may proceed one or more dark-tipped claw-like spurs, the “pedal
spurs.” The terminal joint (tarsus of Pocock) is variously furnished
with hairs and teeth, and always ends in a pair of well-developed
movable claws beneath which a much reduced and sometimes almost obsolete
third claw is distinguishable. The tarsus generally projects in a
“claw-lobe” over the base of the superior claws, and sometimes lateral
lobes are present. The first and second coxae have triangular maxillary
lobes directed towards the mouth. The third and fourth coxae are fused
together on each side, and those on one side are separated from those on
the other by the sternum. In other respects the four pairs of legs are
usually similar.
=Internal Anatomy.=
The =alimentary canal= is a fairly uniform tube, nowhere greatly
dilated. The very small mouth leads into a small suctorial chamber, and
this is connected by a narrow oesophagus, which pierces the cerebral
nerve-mass, with a slightly dilated portion which receives the ducts of
the first pair of gastric glands, often called salivary glands. The
succeeding portion in the prosoma receives four or five more pairs of
ducts from the well-developed gastric glands. In the rapidly narrowing
first metasomatic segment the intestine receives one or two pairs of
Malpighian tubes, and thence proceeds to the anus, situated ventrally in
the last segment.
The =vascular system= is of the usual Arachnid type, the heart being a
seven-chambered dorsal longitudinal vessel lying in a pericardium, with
which it communicates by seven pairs of valvular ostia. Lankester[245]
has demonstrated several pairs of superficial lateral veins connecting
two deep-seated ventral venous trunks with the pericardium. The
lung-books are, so to speak, pushed in to dilatations of these trunks,
so that some of the lateral veins carry blood newly aerated by the
lung-books directly to the pericardium.
The =nervous system= is not greatly concentrated except in the prosoma,
where there is a single ganglionic mass which innervates not only the
whole prosoma but the mesosoma as far as the first and sometimes the
second pair of lung-books. There are two mesosomatic ganglia, variously
situated in different genera, and each metasomatic segment has its
ganglion.
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