The Cambridge natural history, Vol. 04 (of 10)Warburton, Cecil
Science
The Cambridge natural history, Vol. 04 (of 10)
Warburton, Cecil
Animals
=External Structure.=—The external features which the members of this
Order have in common are the segmented pediculate abdomen (9 to 12
segments), the two-jointed non-chelate chelicerae, the antenniform first
pair of legs, and the presence of two pairs of lung-book stigmata
beneath the abdomen. The constituent families differ so much in outward
form that they must be dealt with separately.
The Thelyphonidae or “Whip Scorpions” (see p. 312) have a long-oval
carapace bearing well-developed eyes, two in front, and a group of three
or five on either side some distance behind. The pedipalpi are chelate,
and have their basal joints fused beneath the mouth, being thus
incapable of any masticating motion.
The first legs are six-jointed, and have multi-articulate tarsi; the
others are seven-jointed, and their tarsi, in some species at least, are
tri-articulate. The abdomen consists of two portions, a wide
nine-jointed pre-abdomen and a short narrow three-jointed post-abdomen,
to which a filiform tail is articulated. Beneath the cephalothorax,
between the coxae of the legs, is a distinct sternal plate in two
portions (Fig. 171). The first abdominal ventral plate is largely
developed, and covers two segments. Behind it are the median genital
opening and two pulmonary stigmata, while the other stigmata are behind
the second ventral plate, which corresponds to the third abdominal
segment. On the last abdominal segment there are often two or four
light-coloured spots called “ommatoids,” and considered by some authors
to be organs of sight. Laurie, however (_vide infra_), thinks it more
probable that they are olfactory in function.
The Schizonotidae (see p. 312) have a two-jointed carapace, and do not
possess more than two eyes. There is a short unjointed tail-piece.
In the Tarantulidae (Phrynidae) the whole body is much flattened and
extended laterally, the undivided carapace being reniform, and broader
than long. The long non-chelate pedipalps have their basal joints free
and movable, and there are several sternal plates. There are nine
abdominal tergal plates, the last three diminishing rapidly in size, and
the last plate covering a button-like terminal portion of the abdomen.
The first abdominal ventral plate is largely developed, as in the
Thelyphonidae, and the genital orifice and pulmonary stigmata are in the
same situation as in that group. The Tarantulidae have glutinous glands
in the first abdominal segment which are capable of spinning a few
irregular threads.
In the whole group paired circular depressions are conspicuous dorsally
on all the abdominal segments. These indicate the points of attachment
of the dorso-ventral muscles.
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